Beggiatoa
During his research in Anton de Bary’s laboratory of botany in 1887 , he found that "Beggiatoa" oxidized hydrogen sulfide ( H2S ) as energy source , forming intracellular sulfur droplets , oxygen is the terminal electron acceptor and CO2 is used as carbon source . 
<triplet> CO2 <subj> oxygen <obj> has parts of the class