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messages in their Twitter "home". The following sections are background of the study, research method, results and discussions and finally the conclusion. 2 Background of the study 2.1 Online Social Media trends in disaster Disaster definition by The United Nations Office for Disaster Risk Reduction (UNISDR) is "A serious disruption of the functioning of a community or a society involving widespread human, material, economic or environmental losses and impacts, which exceeds the ability of the affected
of an area gives clues to its potential hazards * Maps, historical accounts, weather and climate data * Rock types, faults, folds, soil composition Geologic Cycle * Geologic conditions govern the type, location, and intensity of natural processes * Sub-cycles * Tectonic cycle * Rock cycle * Hydrologic cycle * Biogeochemical cycles Rock Cycle * Rocks are aggregates of one or more minerals * The rock cycle refers to recycling of three major
everything that goes on, they mistakenly assume that must mean that God is the one causing the disasters as well, when that necessarily is not the case. God allows things to happen, but he doesn't actually force things to happen. Not to say he doesn't arrange some events, maybe he does, but there is generally a more logical reason than that. Sometimes awful stuff just happens. Every now and then disasters are even caused by human irrationality or animosity like the 9/11 attacks, it's easier to play
risks manmade or natural risks (Westall 2002). There are two types of risks manmade and natural disaster. A man-made risk means a disaster to the environment that done by man's actions. This includes, terrorism act, war, automobile or fire risks. It has been said that such risks are highly frequent and less severe in respect to the insurer (Emblemsverg 2008). As a result, a spread of information is available to estimate the likelihood of prospective loss. On the other hand, natural risks are derived
Indian Ocean, thousands of people could have escaped the wrath of the tidal waves that affected a dozen Asian countries in December 2004. Monitoring and advanced warning systems are crucial in reducing the disastrous effects which natural disasters can have as they allow sufficient time to carry out necessary preparations to minimise damage to property and loss of human life. They also permit gas networks, transport systems and nuclear power stations to shut down prior to
However, there continues to be a lack of adequate planning in this area. Physical Impacts The physical impacts of natural disasters include immediate casualties and injuries to persons directly affected by natural disasters as well as secondary health impacts due to damaged infrastructure or loss/contamination of resources. After a natural disaster there may be damage to properties and infrastructures as well as loss of resources needed to survive. This can include loss of crops and safe drinking
difficult part about being a victim of a natural disaster? Part Two: Is Your Community Prepared? Go to www.FEMA.gov and click on one of the disasters that may occur in your community. After learning about this disaster, answer the questions below. 18. What type of disaster did you choose? 19. What preparations does FEMA recommend to avoid loss in this type of disaster? 20. What steps can you take to better prepare your home and family for this type of disaster? Give examples. Your Own Interviews
April-May and October-November (Kuppusamy and Rajarathnam, 2009). Floods Floods are high stream flow that overflows the natural banks of the rivers and most of the times become calamitous. India is the most flood affected nation after Bangladesh. Out of total deaths by Floods in the world, (1/5) are from India. The main causes of floods are excessive rains in river catchments, poor natural drainage, Change of river course, Landslide restricting river flow, cyclone and very intense rainfall (Fig. 1). Over
sea" (Emanuel 453). The system continues to grow larger clouds and increasingly strong winds. The increased amount of moisture in the air and powerful winds feed the storm. Each tropical storm is granted a name. The name helps meteorologists and disaster planners to quickly convey information. Once winds are 74 miles per hour or more, the storm is now classified as a hurricane or its equivalent in another region. By the time the storm becomes a hurricane, it has formed a well-developed eye (see
The largest industrial disaster to date is the Bhopal Disaster. 2)Traffic Accidents: A traffic collision (motor vehicle collision, motor vehicle accident, car accident, or car crash) is when a road vehicle collides with another vehicle, pedestrian, animal, road debris, or other geographical or architectural obstacle causing damage to surroundings which may be severe, injury to health or loss of life. 3)Oil Spill: An oil spill is a release of a liquid petroleum hydrocarbon into the environment
Our first day of providing medical care we saw 147 injured most of which consisted of traumatic amputations typically involving the extremities, large avulsions, compound fractures, lacerations, and severely infected wounds. As Barbara Morita, P.A., one of my fellow CA-6 members, so eloquently put it to words "We saw more severe trauma in a day in Haiti than many clinicians see in a career". During our initial briefing prior to going to Haiti, we as healthcare providers were told that we would
Resources due to Natural Disaster is unavoidable. Therefore, it is necessary to understand loopholes and complexities to change system beforehand to avoid losing of employees. The peer reviewer said that this case study was briefly explained with all scenarios. References Barron, D.L., Phipps V.M. & Steinmeyer P.A. (2005, November 23). After the Disaster 10 Issues for Employers, Transfer of Workers to Other Operations, Retrieved from http://www.workforce.com/articles/after-the-disaster-10-issues-for-employers
After fumes, which contain specific chemicals, rise to upper atmosphere, where the wind is very strong, the harmful gases are spread worldwide. In a chemical reaction, oxides combine with water vapor causing nitric and sulfuric acids as a catalyst using sun light. In the last stage, the acid chemicals in the air are blown into areas where they are mixed into clouds. Moreover, the acids can fall to Earth in three forms: rain, snow, or dry particles (dry deposition). In areas where the weather is dry
by the United Way of Northwest Arkansas. NASW mission is to identify agencies that incorporate the procedures involved in preparedness, mitigation, response and recovery and provide services before, during and after a natural or manmade incident, including weather-related disasters, accidents, terrorism or violent events. NASW utilizes cooperation with Arkansas Department of Health (ADH) and the Arkansas Chapter of the National Association of Social Workers (NASW-AR) the NASW-AR and with the Arkansas
a child should not be denied their right to eat whenever they are in need of nourishment. As a society we should support and encourage the mothers who are contributing to the well-being of our nation by breastfeeding. Breastfeeding is a perfectly natural act of feeding and we should be more supportive of this. Works Cited 10 Benenfits of Breastfeeding. Dir. Gerber. Youtube. Gerber, 8 July 2014. Web. 18 Feb. 2016. "Breastfeeding." Opposing Viewpoints Online Collection. Detroit: Gale, 2016. Opposing
retrospective judgment of nurses' actions in such circumstances is extremely complex. Because of events such as these, ANA ordered a review of altered standards of care to be used during a disaster and in 2007 a conference was held to address those issues.("The free library," December 2006, p. 3) A medical disaster is when the need for care greatly exceeds the available resources. The gap between demand and supply is far worse than a hectic day in the ER and it requires nurse managers to shift their
makes the atmosphere pure. The manufacture of negative ions form soil can help to remove electromagnetic effect so it can help to clear the nature wall and it will not attract dirt. All of advantages of soil can cause people to enjoy living with natural materials. People can decorate the nature walls by making color clay and it make from color sand with mother of pearl or straw that it is enduring and beautiful. Cob buildings prefer soil paint replace using normal coloring. Cob buildings can be suck
had any immediate health effects; however, the signs of radiation induced illnesses can take time to develop (Fukushima…Misunderstood Effects of Radiation). Overall, the health consequences of this disaster will be seen in time, but the economic issues have been immediate. After any natural disaster the cost of cleanup is substantial. The Japanese government is looking at a total economic loss range from $250 to $500 billion US over the next ten years. In order to accommodate this
128 O. Patterson et al. Introduction People make decisions every day, but decision-making in a natural disaster is unique and has great implications for individuals, communities and society at large. Hurricane Katrina clearly demonstrated the need for sound planning, swift execution and efficient accountability; but ultimately the negative consequences people experienced in this (as in any) disaster were conditioned by their perceptions of risk and their vulnerability, and how these factors influenced
Stillman, D., 1982) The components of this disaster were not purely negative, but when looked at through the eyes of an outsider, the negatives outweighed the positives. Though many factors made up this event, health was a huge concern in the history of this disaster. Among the people of the twenty-two villages that were evacuated, individuals were concerned because of mysterious illnesses occurring. (Soran, D. & Stillman, D., January 1982) Victims of the disaster were seen with the skin of their faces
Although the reason for the Chernobyl disaster occurred long before the accident happened, the construction and implementation of the reactors is not the only factor involved with the meltdown. The Russians operated on the RMBK-1000 series reactor; four of them were stationed at Chernobyl. Rector number 4 was scheduled to be shut down to allow for routine maintenance the day before. Taking advantage of the shutdown, the plant operators on duty performed a test with the reactor's residual kinetic
CERTs use ICS to manage the emergency response operations. The Incident Commander is the leader of the entire disaster relief efforts. Following the Incident Commander, the Operations Section Chief, Logistics Section Chief, Planning Section Chief, and Administration Section Chief all have specific specialized duties that aid in the disaster relief efforts. Depending on the situation, the ICS undergoes a metamorphosis to handle the needs of the emergency. Within the command ladder two-way communication
location at its top speed, arriving two hours after the Titanic submerged. All but one of the twenty lifeboats was found. A month after the collision, the last of the Titanic's lifeboat was founded, but there were three dead bodies inside of it. The disaster drastically caught the public's attention worldwide. As a result of the reactions of many, governments of nations
implementation performed in relation to the strategy formulation's needs and findings. The strategic evaluation is important in allowing for the strategic implementation to be one that has satisfied the needs of the strategic formulation. BAA Terminal Disaster BAA opened Heathrow's fifth terminal for business on 27th March, after 6 years of
Group appointed to monitor the leakage flow rates. According to an estimate, the leaked amount is about 4.9 million barrels of oil. It is also undermined the 1979 Ixtoc-1 oil spill which was the largest spill on record before this Deepwater Horizon disaster in the Gulf of Mexico. Three fundamental strategies for removing and cleaning of spilled oil were designed. First, contain it on the; second, to dilute and disperse the oil in less sensitive areas; and third, to remove the oil from the water. These
damages to the "public's natural resources" (McCammon 2003), their overall behaviour in dealing with compensation has been questionable. On the one hand, their dealings with the US Federal government indicate that they are complying with all governmental laws, rules and regulations applicable to its business (ExxonMobil n.d.) and have indeed met their legal obligations. Their settlement with the US Federal government included a civil settlement of $900 million to restore the natural resources, criminal
Several factors that occurred on that day and the previous days might have contributed to the disaster. The tank was constructed poorly and tested insufficiently. Due to fermentation occurring within the tank, carbon dioxide production might have raised the internal pressure. The rise in local temperatures that occurred over the previous day also would have assisted in building this pressure. Records show that the air temperature rose from 2°F to 41°F (from −17°C to 5°C) over that period. The failure
taking place outside of the typical office. The counseling will hold a session in a home, community, or where the client is staying for the time being. The strengths based approach of the CCP recognizes that although the client is suffering in a disaster, the client is a survivor. The client is asked to reflect upon other circumstances in which hardship has been overcome and what hardships are being overcome in the present moment. The client is asked what can be learned from the current adversity
in the south western area, natural hazards increase the vulnerability of the coastal dwellers and slow down the process of social and economic development. Significant country features include: • A vast network of rivers and channels • An enormous discharge of water heavily laden with sediments • A large number of islands in between the channels • A shallow northern Bay of Bengal and funneling to the coastal area of Bangladesh • Strong tidal and wind action Natural and human induced hazards
People search for family through destroyed houses at Sirombu village in Nias, North Sumatra, Indonesia, Dec. 29, 2004. Not only was Indonesia hit dramatically by this disaster, other areas included: Somalia where approximately 300 were said to be killed, Maldives where the death toll was 82, the tsunami left 37 of the Maldives's roughly 200 inhabited islands temporarily unsuitable for human habitation, destroying drinking water and food supplies as well as communications and power generation


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HSC/ HSLC Exam Time Table And HSC/ HSLC Exam Date Sheet 2017 here are now available online. after that you all students also can check it from here.
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Students who are appearing from the number of different schools in the 10th 12th Board Exam 2017 for this session year 2016-17 can download the 10th 12th Time Table 2017 for all different admitted state and Center Board on the official Examination and Time Table link of Board. Every year these each State and Central Board have to conducts Class Xth and Class Xllth Exam in the month of March/April. So this running Academic Session year the Boards will be also conduct the HSC Examination 2017 for academic year 2016-17. Here we are also providing all the HSC Exam Date sheet 2017 such as SSLC/HSSC Time Table 2017, SSC/HSC Exam Schedule 2017, High School/Higher Secondary Exam Date Sheet 2017, Inter Class Date Sheet 2017, Matric/Intermediate Exam Schedule 2017, 10th 12th Exam Time Table 2017 etc.
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text: All Courses Applying to Hull Accommodation Money Visit us Taught MSc in Clinical Anatomy and Education How to applyMedicine About the course The MSc in Clinical Anatomy and education is designed to provide advanced training in clinically applied human anatomy. The programme is built around core dissection-based modules and offers you the opportunity to conduct whole body dissection in a newly developed Thiel embalmed and plastination facility. You can tailor the programme to suit your individual interests and will learn from expert tutors as well as experienced clinicians. This programme is designed especially for intercalated medical students, anatomy teachers, trainee surgeons, physiotherapists, sports scientists and other health professionals. In addition to developing advanced anatomical skills and knowledge, this programme provides a comprehensive professional education that focuses on developing teachers and researchers of anatomy. You will be trained in pedagogical techniques that will allow you to teach anatomy to students and health care professionals. Key Facts Tuition Fees UK/EU: £7,000 full-time / £3,500 part-time (2 years) / £2,340 part-time (3 years) A new system of postgraduate loans for Masters degrees in the UK are now available. They provide up to £10,280 for full-time and part-time taught and research Masters courses in all subject areas. For more information, visit our Postgraduate Loans page. Overseas: £20,100 full-time / £10,050 part-time (2 years) / £6,700 part-time (3 years) Scholarships are available for this programme, to find out more, please visit: www.hyms.ac.uk/clinical-anatomy-and-education. More information on fees can be found in the funding information section of this page. Application deadlines To apply, or for more information, please visit www.hyms.ac.uk/postgraduate/how-to-apply/applying-for-postgraduate-study. Entry requirements Minimum Applicants to the MSc in Clinical Anatomy / Clinical Anatomy with Education must have previous training in human or mammalian biology. (e.g. graduates from medical, biomedical, anatomical science, veterinary medicine, and zoology backgrounds). Intercalating students must have successfully completed of a minimum of three years of an MB BS or comparable medical qualification. International students Please see the HYMS website for details Admissions status Open for admission in 2017/18 Mode of study: All core modules are delivered at the University of Hull. Some electives are available online or at the University of York. Duration: 1 year full-time / 2-3 years part-time Tutor details firstname.lastname@example.com +44 (0)1904 321782 Hull York Medical School Research International enquiries Related courses MSc in Human Anatomy and Evolution MSc in Health Professions Education (full-time) MSc in Translational Oncology MSc in Colonoscopy MSc in Cancer Imaging Medicine research degrees MSc in Clinical Anatomy MSc in Physician Associate Studies Certificate/Diploma/MSc in Health Professions Education Accommodation From state-of-the-art halls to nearby student suburbs Study information This programme offers a mix of core and elective modules, giving you the opportunity to develop fundamental anatomical knowledge whilst also allowing you to specialise in specific topics suited to your professional interests. Due to the education element of this programme, two of your electives should be education modules. Core modules include: Clinical anatomy of the limbs and spine Clinical anatomy of the trunk Clinical anatomy of the head, neck and brain Research Project/Dissertation Education Electives: Learning and Teaching Assessment and Feedback Contemporary Issues in Health Professions Education Research Electives: Research Methods & Statistics module Research Approaches in Health Professions Education module Anatomy Electives: Tissues of the Body Hard Tissue Biology Human Evolutionary Anatomy * All modules are subject to availability. Funding information Graduate Studentship The University of Hull is pleased to offer graduates progressing from undergraduate to postgraduate taught study a studentship towards the cost of their tuition fees. Find out if you're eligible by visiting University of Hull Graduate Studentship webpage. International Scholarships and Bursaries For a list of all scholarships and bursaries for international students, please visit the International Scholarships and Bursaries webpage. UK and EU Scholarships and Bursaries Please download our Funding Your Postgraduate Studies document for in-depth guidance on funding sources. For more information, please visit the UK and EU Scholarships and Bursaries webpage. Future prospects This programme will enable you to develop advanced anatomical skills and knowledge through the application of topographical, biomechanical and educational knowledge to translate the experience of handling cadaveric anatomy to the functional anatomy of the living person. You will learn to critically appraise and reflectively apply advanced anatomical acumen to inform clinical decision making. The programme will develop your knowledge and skills to undertake and deliver high quality research in clinical anatomy and related disciplines. 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Gold Rush was the term used to describe the economic boom that resulted from the discovery and subsequent exploitation of Imulsion following the Era of Silence, and was a major turning point in Sera's history. With the discovery of Imulsion and the invention of the Lightmass Process, an entirely new industry quickly grew into being and supplied thousands of new jobs, even as the planet's energy problems seem to disappear with an influx of this new, cheap, and abundant source of energy. However, a few problems came from this. Those involved in the immediate extraction and processing of Imulsion suffered a variety of health complications and a shortened life span as a result of exposure to the substance, though this was largely considered a small price to pay for Sera's energy security. These health concerns would eventually reappear nearly a century later in the form of Rust Lung.[1] Eventually, however, rampant speculation on Imulsion and the ever-increasing demand for the substance led to a major financial collapse, and eventually the Pendulum Wars, ending the Gold Rush.[2][3]


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Twins third baseman Danny Valencia got the best of longtime friend and Indians closer Chris Perez Tuesday night in Minneapolis.
Trailing 1-0 in the bottom of the ninth, Valencia hit a walk-off single with the bases loaded to give the Twins a 2-1 victory.
Less than an hour later, Perez met a smiling Valencia, who had invited Perez over to his house for the night.
It was not the welcome Perez was looking for.
The two have been friends since high school and were teammates at the University of Miami, where they were roommates.
"It's just nice - it's huge - to be able to compete with one of your friends, let alone come up with a huge knock against them," Valencia said after the game.
Valenica hit a fastball that fell in and broke his bat.
"At least I got the bat," Perez said. "He can't be happy about that hit, just he's happy because they won. I'll take that swing every time, just fell in there."
Valencia struck out against Perez in his only other at-bat against him in 2010.
It was just the second blown save all season for Perez. An Indians win would have given them an eight-game lead over the Twins in the American League Central, instead the Twins moved within six games.
The Twins close out the four-game series today.
Asked what the postgame plans were for the evening? "We're just going to watch the highlight over and over again," Valencia said laughing.
Daily Pitch is a baseball fan's destination for breaking news, emerging trends and enlightening banter. With a veteran group of reporters positioned in three time zones, we provide a balanced, comprehensive glimpse at our national pastime, with a nod to both the passion and complexities that make the sport irresistible to many. More about our reporters


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Koala in a Gum Leaf Bed
Approximate weight including packaging 0.02kg
This tiny koala is rather shy and likes to hide inside his gum leaf bed.
The koala and gum leaf are made from wool-blend felt and hand-sewn with fine attention to detail. No glue involved, just lots of tiny stitches.
Hand-cut. Hand-sewn. Made in South Australia by a local artist.
The koala is 4cm x 3cm. The gum leaf bed is 13cm x 5cm.
Please Note: This item may not be suitable for children under three years of age due to the use of beads and the small size of some components.
The colour design and size of all items remain the same, but the stitching may vary slightly from the photo shown. This is because each item is sewn with great attention to detail.


text: The Crawlspace is the space in the Aqua Teen residence where they go to the bathroom, (since they don't have one) and discover some things. The crawlspace in "Universal Remonster" AppearancesEdit * Love Mummy * Super Hero * Universal Remonster' * Rubberman Retrieved from "http://aqua-teen-hunger-force.wikia.com/wiki/Crawlspace?oldid=9459" Ad blocker interference detected! Wikia is a free-to-use site that makes money from advertising. We have a modified experience for viewers using ad blockers Wikia is not accessible if you've made further modifications. Remove the custom ad blocker rule(s) and the page will load as expected. Categories: Locations Add category Cancel Save


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Bookmark/Search this post with
Rugby League field, Chairmans Lounge , Heritage Lounge, Community Lounge, Grandstands and sporting infrastructure, Village green, Cricket nets, Sportsfield lighting
Public toilets, Playground, Parking, Disabled Toilets, Bus Stop within 100m, Enclosed Playground, Kiosk, Garbage Bin
Brookvale Park includes Brookvale Oval and an open park (the village green) which are managed and maintained for the community by Warringah Council.
Affectionately known as "Fortress Brookie", Brookvale Oval is home to the Sea Eagles Rugby League team and the Brookvale Show.
You can book the Oval or any of the three community rooms.


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About the Book
Composed for my senior project, this is the collection of my favorite recipes ranging from healthy breakfasts to decadent desserts and every mean in between.
Features & Details
- Category Cooking
Standard Portrait, 7.75×9.75 in, 20×25 cm
82 Pages
- Publish Date Mar 18, 2014
- Language English
- Tags Cooking, Baking, Food, Vegan, Diets, Teen, High-School, All-purpose
See More
About the Creator
Tori Lorvig
New Hamshire
Hi my name's Tori and I'm a senior in a small town in NH. I made both of my cookbooks as part of my graduation requirement. I've loved cooking since my toddling years and am delighted to be able to share my recipes with you.
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We've farmed organic since the day we planted our first heirloom tomatoes over 30 years ago. It's good for the land, and it's great for you.
There's a lot of buzz about sustainable farming, and we couldn't be happier. We've seen how much richer the soil is when we control weeds, pests and disease without toxic chemicals. We're pretty sure it makes our harvest taste better, too.
When we started to farm, it was just the two of us. Now we've got kids of our own and 10 employees who've become our extended family. We're all here because we absolutely love what we do.


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House Party OrgyFavorites [+]
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You can now enjoy this movie on your iPhone®, iPad®, or compatible Smart TV & Android devices.
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Available MP4 sizes:SD 480p (1.47 GB)
DRM-free downloads offer content that is not limited by standard Digital Rights Management (DRM) access technologies. These technologies allow hardware manufacturers, publishers, copyright holders and individuals to limit the usage of digital content.
When you purchase DRM-free downloadable content, that content is available for you to download on any computer and save to a disk whenever you like. You can also download to multiple computers. There is no timeframe associated with your purchase and the content will not expire.
Running Time: 94 minutes
Released: 06/2013
Studio Name: Zero Tolerance
Director: Mike Quasar
Description: Never mind the chips and dip, this party is your ultimate fantasy! Smokin' hot girls everywhere you look and no need to find a separate room to get naked! Join the shindig that's busting with horny babes who crave naughty group sex, trying out every position imaginable with everyone in the room! You won't be sorry you came!
2 Scenes from House Party Orgy
House Party Orgy: Scene 1 Free Preview
Starring: Courtney Taylor Eve Evans ... More Details
Stars: Courtney Taylor, Eve Evans, Alison Tyler, Cali Carter, Riley Reece
Sex Acts: Breast Play, Facesitting, Rim Job, Orgy, Girl on Girl action, Masturbation, Oral Sex
Settings: Living Room
Studio: Zero Tolerance
House Party Orgy: Scene 2 Free Preview
Starring: John Strong Courtney Taylor ... More Details
Stars: John Strong, Courtney Taylor, Tommy Gunn, Marcus London, Anthony Rosano, Tommy Pistol, Eve Evans, Alison Tyler, Cali Carter, Riley Reece
Sex Acts: Breast Play, Rim Job, Orgy, Handjob, Masturbation, Vaginal Sex, Deep Throating, Oral Sex
Positions: Standing Side Saddle, Standing Doggy Style, Standing, Standing Missionary, Side Saddle, Missionary, Laying, Reverse Cowgirl, Cowgirl
Settings: Living Room
Studio: Zero Tolerance
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I was playing at Sweetwater Wetlands and focused on the reflection instead of the sparrow himself. So I flipped the image. Thought it was fun and wanted to share~☺
Tucson, Arizona
Canon Powershot sx10is
White-crowned Sparrow
This hardy sparrow is found in many habitats, from muskeg and recent burns in boreal forests to coastal sage scrub in California; migrants and wintering birds are found in brushy habitats.
Scratches feet along the ground, foraging for insects, caterpillars, and seeds. Also gleans food from vegetation.
VOICE Song is usually a series of scratchy whistled notes, one higher, one lower. Call a flat, low pink or chink , sharper and more ringing than Golden-crowned.


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All three versions are equipped with Dolby Atmos, and a free DTS:X update is due later in 2016. After the update, owners will also be able to deploy Dolby Surround and DTS Neural:X upmixers to create 3D sound from legacy Dolby or DTS soundtracks.
Bluetooth, wi-fi, Apple AirPlay and Spotify Connect compatibility are standard across the range, and there's support for high-resolution audio files including FLAC, ALAC, AIFF, WAV and DSD (up to 5.6MHz).
Video is equally well catered for, with the latest HDMI 2.0a spec, as well as HDCP 2.2 on all inputs, required to handle copy-protected 4K Ultra HD content.
We are testing the Denon AVR-X2300W right now
All three are 7.2-channel amps with power ratings of 145W, 150W and 180W respectively. The AVR-X1300W can deliver Atmos in a 5.1.2 configuration and has six HDMI inputs.
Additional features on the AVR-X2300W include upgraded sound tuning, eight HDMI inputs, advanced video processing with 4K upscaling for HDMI sources plus ISF (Imaging Science Foundation) video calibration. And the top-of-the-range AVR-X330W gets analogue-to-HDMI conversion, advanced multi-room options, a 7.2 channel preout and RS232 control.
More after the break
There's plenty of connectivity, even on the entry-level AVR-X1300W
An upgraded version of Denon's AVR remote app is available for iOS and Android devices as well as the Kindle Fire.
The Denon AVR-X1300W and X2300W are available now, with the X3300W due to arrive in August. We already have the AVR-X2300W in our test rooms and will publish the official review online later this week.


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Taking the stage to present Lohan with Movieline's Hollywood Life Superstar Award, Ann-Margret, 64, grasped her microphone and bent it toward her mouth. "My husband always says you've got to pull this toward your ... ," she began before stopping herself midsentence when the dirty-minded audience erupted into laughter.
"I forgot what a young group this is," the embarrassed star continued.
With sassy Ann-Margret as her role model, it is no wonder that Lohan also has her own momentary flirtations with controversy. But in the lobby before the show, the newly blond Lohan was on her best behavior. She squatted down to sign a fan's jeans and said, "The fact that Ann-Margret came here to be with me is incredible."
Looking very Lohan herself was red-haired Desperate Housewives star Marcia Cross, now so confident with her heterosexuality that she planted a kiss on fellow housewife Felicity Huffman at the previous evening's GLAAD (Gay and Lesbian Alliance Against Defamation) Awards. "I thought the kiss was a little short," laughed a beaming Cross, who came to the Young Hollywood event dressed in a sleek, silvery white ensemble.
"I was just getting warmed up, and boom! It was over."
Cross was on hand to present co-star Jesse Metcalfe with the Exciting New Face award. "The old face just wasn't doing it for me," Metcalfe joked. "I'm glad people think this new one's exciting."
Half of this summer's Fantastic Four superhero quartet — Jessica Alba and Chris Evans — were named male and female Superstars of Tomorrow. Seated in the audience, Evans howled when show host Aisha Tyler made a reference to the bone-thin women of WB. See, one of those WB skinny minnies is Evans' sometimes girlfriend Jessica Biel (7th Heaven), whom Evans implied at the after-party is not his steady.
Stefani attended with husband Gavin Rossdale, who received the Standout Performance award for his role in Constantine.
Asked what makes him a standout husband, Rossdale said, "You have to ask Gwen that." (She wasn't talking.) He said they'll be dining Sunday with Stefani's mother for Mother's Day and will "keep practicing" to perhaps be parents themselves in 2006. "You never know ... "
Also standing by her man: Bo Derek, whose beau, John Corbett, was a presenter. She said the secret to their success as a couple is simply "being in love." Derek said she'd like to get married again but let it be known that she's "a very happy independent person."


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Packaging & Bagging
Flexicon Launches 250-Page Spanish WebsiteOctober 5, 2015
Flexicon has launched an all-new 250-page Spanish language website detailing the company's full line of bulk handling equipment from stand-alone units to integrated plant-wide systems. The new site, www.flexicon.es, coincides with the 2015 opening of the company office...comment
Industrial Vacuum for Continuous BaggingSeptember 29, 2015
The Model 850 industrial vacuum cleaner is ideal for applications requiring the clean-up of dry, dusty debris, ranging from fine powders up to 2-in. solids. Bulk materials are disposed of in a safe, dust-free, and convenient manner via a continuous poly-bagging system, eliminating dumping of a drum...comment
Calcium-Free Polypropylene Fabric Eliminates DegradationSeptember 25, 2015
Calcium carbonate (CaCO3) is one of the most popular mineral fillers used in the plastics industry. Its wide availability makes it an economical component in the production of polypropylene bags. CaCO3 helps as a whitener, lubricant, and finishing agent in the manufacturing of FIBCs. Used in the...comment
World Packaging Machinery Demand to Reach $52.5 BillionSeptember 25, 2015
Global demand for packaging machinery is projected to advance 6.5 percent per annum through 2019 to $52.5 billion. Product sales will be driven by market growth in China and other industrializing nations, where demand for processed food and beverage products and other manufactured goods is expected...comment
Haver Filling Systems Now Haver & Boecker USASeptember 17, 2015
Haver Filling Systems, a provider of solutions in material handling, bag packaging, liquid and pasty filling equipment, and palletizing systems, is changing its name to Haver & Boecker USA. "Haver & Boecker USA's focus from the beginning has been to offer our customers with...comment
Filling System Mines Competitive Edge for Kaolin ProducerSeptember 14, 2015
The sunny state of Georgia is famous for peaches, pecans, and southern hospitality, but Georgia is also famous for kaolin. Commonly known as china clay, kaolin is an important mineral used around the world in hundreds of products such as paper, porcelain, paints, sealants, adhesives, plastics,...comment
Rise & Fall Bulk Bag FillerSeptember 11, 2015
The Spirofil rise & fall bulk bag filler was designed specifically to handle fragile fruits and nuts. It rises to just below the outlet of a filling tube filling bags with whole nuts, while providing a soft landing for the fragile products. As the bag fills, it gently lowers, or falls, to...comment
Iris Diaphragm Valve for Big Bag Discharge StationsSeptember 11, 2015
The Mucon JS iris diaphragm valve is specifically designed for fitting to big bag discharge stations for the controlled emptying of product. Available in three sizes – 12-, 15-, and 18-in.-diam – the valve is suitable for dry powder materials up to 62 lb/cu ft. The single diaphragm and...comment
The Bulk Bag Companycomment
Bulk Bag Weigh FillersAugust 31, 2015
Spiroflow Systems provides accurate gain-in-weight bulk bag filling for low- to medium-volume end-users with the Spirofil C Series bulk bag weigh fillers. Up to 20 bulk bags can be filled per hour. Standard accuracy is 1% or better, with typical accuracies of +/-.5%. Standard features of this...comment


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Dr. Carolyn Kurylas MD Patient Reviews
- Anesthesiology Phoenix, AZ
Overall Rating
7 Patient Reviews
- Wait Time 5 minutes
- Appropriate Followup
- Spends Time With Patients
- Bedside Manner
- Easy Appointments
- Friendly Staff
- Promptness
- Fair and Accurate Diagnosis
Showing 1 - 7 reviews
1.0 of 5
February 10th, 2015
greedy doctor
went in for a colonoscopy that is supposed to be covered %100 by my insurance. My doctor is in network and so is the hospital I went to. Dr Kurylas shows up while I am waiting in pre-op and says she will be the anestisiologist. Turns out she is not in network and now will not accept the amount the insurance company pays.. She is demanding that I pay her an additional $800 for her services that I never hired her to perform. Never would recommend this doctor. This is so wrong.
1.0 of 5
April 10th, 2012
bad experience
I experienced a much longer recovery time from anesthesia administered by this doctor than for any previous surgery. Her billing staff is in disarray and has hounded me for a bill I paid months ago, causing me to waste time and energy on trying to square my account. For a subsequent surgery I asked that she not be involved in any way. Am investigating lodging a formal complaint against her.
5.0 of 5
April 21st, 2011
The Very BEST
After 72 years I was given the greatest anesthesia by Dr. Kurylas. She caring, attentive, reassuring and made my surgery easy. With many health issues she was knowledgable and allowed me and my family the peace we needed. She was not only delightful as a person she was extremely learned in anesthesia. I had no issues after surgery with nausea as I usually do. What a great experience.
1.0 of 5
January 19th, 2011
This woman is a wreck
she takes her license to bill however much she wants very seriously. never seen such an inflated bill for 10 minutes of work no matter what the complexity.
5.0 of 5
May 14th, 2015
3.7 of 5
January 30th, 2015
5.0 of 5
October 24th, 2011
Health Guides
Type 2 Diabetes
Type 2 diabetes is a chronic disease in which your body is unable to maintain a normal blood sugar (glucose) level.
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Dr. Carolyn Kurylas, MD is similar to these 3 Doctors near Phoenix, AZ.
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A Breakfast Invitation
Created November 2012 (Third Edition)
This is an invitation to enjoy the most important meal of the day-Breakfast!
Cover Photo: Perfect Scrambled Eggs
Lovingly Compiled by
Gail Welch [sugarshack]
Get Your Copy!
Order a professionally printed, perfect bound cookbook with a full color glossy cover and 50 recipes.
Or Read It Now On Your...


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Product Spotlight
Featured Research
Quick Stats
You are here
Knowledge Is Power
A well-crafted digital marketing strategy can help grocers engage meaningfully with produce shoppers.
According to a study by Deloitte Digital, digital technologies influence 36 percent, or $1.1 trillion, of in-store retail sales, which will likely increase to 50 percent by the end of 2014. Eighty-four percent of visitors reported using digital for shopping-related activities before or during their most recent trip to a store, and 22 percent of consumers spend more as a result of using digital.
What makes digitally savvy consumers tick, and why do they buy? Answering these questions is becoming increasingly critical to your store?s bottom line.
A New Breed of Shoppers
Shoppers increasingly rely on digital tools to share information with one another and to create ?two-way? conversations with brands and companies. These people are more informed, more engaged, more opinionated and expect more from brands of all kinds. And because they?re quick to share their opinions, they can be a tremendous source of insight to your company and your most powerful advocate ? or your loudest protester. When PMA worked on updating its strategic plan last year, the effects of increased involvement from the digitally connected consumer surfaced among members? chief concerns for the future. That?s why June?s PMA Fresh Connections: Retail event centers on the digitally savvy consumer.
You might think these digitally savvy shoppers are making your job more complicated, and on some level that?s true. But more importantly, there?s tremendous opportunity for grocery retailers to use digital media, along with a deeper understanding of connected consumers, to eliminate barriers and build consumption of fresh fruits and vegetables. To do so, however, everyone along the produce supply chain needs to understand ever-evolving technology and the new ways shoppers make decisions.
Consumer Demand Speaks
Doug Madenberg and Brian Numainville, both principals at the Lake Success, N.Y.-based Retail Feedback Group and speakers at PMA Fresh Connections: Retail, have conducted numerous studies on consumers? online shopping behaviors. They find digital grocery technology plays out differently for individual retailers in different regions. They also find success comes from knowing how to optimize your digital strategy to target communications based on customer type, economics, competitive landscape, and other factors unique to your store. Supermarket retailers need to learn more about advances in online shopping, since failure to do so puts their stores at risk of falling behind on responding to a growing consumer interest.
Along with the rise of the digitally connected world comes a whole new set of variables for understanding the consumer. Another part of the PMA Fresh Connections: Retail program explores ?psychographic segmentation.? This type of consumer demarcation looks at personal data such as values, interests, activities, opinions and lifestyles, rather than demographics, to uncover consumers? needs and motivations, both articulated and unarticulated.
The presenters, Steve Longley, CEO of Philadelphia-based TPG Direct, and Brent Walker, VP and CMO of C2B Solutions, in Mason, Ohio, say the advantage of psychographic segmentation is that its results are more effective in resonating with a consumer?s motivations and unspoken needs than traditional ways of segmenting consumers, like demographics, deliver. Psychographic segmentation is helping many retailers understand the deeper motives that drive an individual?s shopping behavior, which allows companies to adapt to the needs, wants and values of their target audience and gain insight into purchasing drivers, risk adversity and relationship styles.
The More You Know
Part I of Progressive Grocer Independent?s ?Technology and the Independent Grocer? research study focuses on marketing and digital technologies, including online, social and mobile technology. The study finds independent retailers identify sustained engagement, messaging relevance and staying on top of current trends in social media among their greatest issues when it comes to connecting with shoppers today. As the report points out, ?? the challenge in addressing this market isn?t a lack of digital marketing tools; on the contrary, it?s an overabundance of them.?
The same holds true for grocery retailers of all sizes. No doubt, digital and social media technology move at a dizzying rate. The best defense against not getting lost in all this is to seek industry-specific resources on these topics, such as PMA Fresh Connections: Retail or the new PMA website (www.pma.com), both of which can keep you apprised of the digital advancements and the connected consumer. The new site has a search function that knows what you?re interested in based on your browsing history, and delivers customized content in tune with your interests. So if you?re exploring topics such as digital consumer or social media, you?ll see related articles and information appear on the right side of your screen. Presentations from PMA Fresh Connections: Retail will be posted on the site in the coming weeks.
After perusing this information, use everything you learn to get reacquainted with your shoppers at the deeper levels necessary to be meaningful to them in today?s digital environment. When you discover where your shoppers? values, opinions and technology platforms intersect with your brand, you?ll have a digital marketing foundation to provide a two-way dialog for delivering what shoppers are interested in and for gaining insight that you can feed back into your digital strategy to further build engagement, loyalty and sales.
There?s tremendous opportunity for grocery retailers to use digital media ? to eliminate barriers and build consumption of fresh fruits and vegetables.


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Tuberculosis, which was once a fairly common condition, is now considered to be a "rare disease" by the National Organization of Rare Diseases. A bacterial infection that affects the lungs, kidney, respiratory system and the spine, tuberculosis can be deadly. This condition has a variety of different names, however, so it's important to know the aliases that tuberculosis goes by.
Perhaps the most common name of tuberculosis was consumption. This name was associated with the eventual deterioration of the person's health, as well as his lungs.
The obvious choice of nickname, tuberculosis is often shortened to simply TB. This name is used when a person is tested, simply called a TB test.
Koch's Disease
Robert Koch was the German doctor who discovered the bacteria that causes tuberculosis. As such, it's sometimes referred to as Koch's disease.
Wasting Disease
This name refers to the fact that a person who has tuberculosis begins to waste away. The symptoms of tuberculosis first steal a person's energy, and then her physical mass as she "wastes" or is "consumed."
White Plague
Another name for tuberculosis, though not one that's used often anymore, is "white plague"--white refers to the victims' loss of skin color and plague refers to the high death rate.
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Tuberculosis is a disease caused by the bacteria Mycobacterium tuberculosis. Although this disease most commonly infects the pulmonary system (the lungs), about...
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Leaderless genes in bacteria: clue to the evolution of translation initiation mechanisms in prokaryotes
Shine-Dalgarno (SD) signal has long been viewed as the dominant translation initiation signal in prokaryotes. Recently, leaderless genes, which lack 5'-untranslated regions (5'-UTR) on their mRNAs, have been shown abundant in archaea. However, current large-scale
Here, we study signals in translation initiation regions of all genes over 953 bacterial and 72 archaeal genomes, then make an effort to construct an evolutionary scenario in view of leaderless genes in bacteria. With an algorithm designed to identify multi-signal in upstream regions of genes for a genome, we classify all genes into SD-led, TA-led and atypical genes according to the category of the most probable signal in their upstream sequences. Particularly, occurrence of TA-like signals about 10 bp upstream to translation initiation site (TIS) in bacteria most probably means leaderless genes.
Our analysis reveals that leaderless genes are totally widespread, although not dominant, in a variety of bacteria. Especially for
As the first stage of protein synthesis in gene expression, translation is a key process highly conserved in the biological system. Up to now, 31 universally occurring genes identified in 191 species are shown being involved in the translation process
Regarding the diversity of translation initiation mechanism, SD initiation has long been considered the dominant way in prokaryotes. However, recent studies revealed that leaderless initiation should be as important as SD initiation in archaea, one of two branches in prokaryotes. Computational study of 144 genes in the archaeal
In spite of above-noted efforts to understand on archaea, little is known about the bacteria-wide situation of leaderless genes. One of major reasons is that translation initiation was believed more complex in archaea than in bacteria
The objective of this study is to answer the question to understand the translation initiation mechanism and its evolutionary scenario in view of leaderless genes in bacteria. We developed an algorithm, which is validated with statistical significance, to classify the initiation regulatory signals upstream to gene start into SD-like, TA-like and atypical signals for all genes in each prokaryotic genome. The method leads to definite identification of both leadered and leaderless genes in a genome. We examined 953 bacterial and 72 archaeal genomes and annotated diverse translation initiation signals in these genomes. Focusing on the leaderless genes and their initiation signals, our analysis reveals that leaderless genes are totally widespread, although not dominant, in a variety of groups in bacteria. What more useful is the quantitative relationships of evolution of initiation signals in bacteria, which might provide a clear picture of the evolution of translation initiation mechanisms.
We have performed a thorough analysis on 953 bacterial as well as 72 archaeal genomes (TIS upstream sequences 20 bp for bacteria and 50 bp for archaea) by classifying all genes in each genome into categories of being SD-led, TA-led or atypical, meanwhile annotating the translation initiation signal for each gene. The definition and algorithm can be seen in
Figure 1
Sequence logos of TA-like signals in bacteria
Sequence logos of TA-like signals in bacteria. Sequence logo
Figure 2
Positional distribution of TA-like signals to translation initiation site (TIS) in bacteria
Positional distribution of TA-like signals to translation initiation site (TIS) in bacteria. Four bacterial genomes are:
Validation of the algorithm
To demonstrate how sure we can be that the relationship between the signals detected through the algorithm and gene translational initiation, we first show evidence for the statistical significance of the TA-like signals found in bacterial genomes, and then estimate the accuracy of our prediction for leaderless genes. A limiting case of examples is the
Figure 3
Validation of the signal detection algorithm
Validation of the signal detection algorithm. (A) Distribution of number of TA-like signals identified in randomly shuffled TIS upstream sequences retaining dinucleotide frequency of 7769 genes for the
A search is further performed in Google Scholar with the key words "leaderless and
Table 1
Documented leaderless genes in
Gene name
Sequence upstream of TIS1
1The last three characters are triplet for start codons, experimentally determined TSSs are bold, and the most-likely Pribnow boxs are italic. The -10 promoters proposed in literature, if available, are underlined.
2The probability that a gene is a leaderless gene regulated by σ70 was calculated by Eq. 5.
It should be noted that the current algorithm is not sensitive enough to the genome when it has a few leaderless genes. In addition, with a widely-used Pribnow box as reference consensus (see
High usage of leaderless genes in archaea
Based on the leaderless genes identified for each genome, we then present an overall view of leaderless genes in all archaea studied here (see Table
Table 2
Distribution of leaderless genes in sequenced prokaryotic genomes
Avg. Percentage2 (%)
21 (/24)
49.5 (6.1, 66.7)
36 (/46)
31.7 (3.8, 63.3)
1 (/1)
71.8 (71.8, 71.8)
Other Archaea
1 (/1)
41.2 (41.2,41.2)
3 (/3)
13.3 (9.7, 19.4)
86 (/89)
19.2 (2.3, 29.8)
2 (/5)
26.5 (20.7, 32.4)
1 (/37)
2.4 (2.4, 2.4)
4 (/12)
4.7 (4.0, 5.7)
5 (/5)
39.4 (35.9, 46.1)
56 (/209)
4.2 (2.4, 16.6)
2 (/117)
6.3 (4.7, 7.8)
9 (/71)
6.8 (2.8, 10.6)
15 (/239)
4.5 (3.0, 5.7)
8 (/33)
15.8 (5.7, 20.6)
2 (/26)
2.9 (2.7, 3.2)
6 (/18)
3.9 (1.8, 6.9)
Other Bacteria
8 (/20)
7.8 (1.4, 16.1)
1 The number of genomes having leaderless genes. The numbers in parentheses show how many genomes have been sequenced in these groups.
2 The average percentages of leaderless genes. The numbers in brackets are the minimum and maximum percentages of leaderless genes in these genomes.
Additional file 1
All Latin names mentioned in the manuscript and their taxonomic relations. This additional file is a table listing all Latin names mentioned in the manuscript and their taxonomic relations.
Click here for file
Diverse distribution of leaderless genes in bacteria
It is of great interest in this study to discuss the leaderless genes in bacteria. Below we report the analysis of leaderless genes in all bacterial genomes (see Table
It has been reported that metabolic-related genes, especially for energy production and conversion, tend to show a higher proportion of SD-containing genes
Figure 4
Relative proportion of leaderless genes in COG function categories
Relative proportion of leaderless genes in COG function categories. The function categories are: A, RNA processing and modification (information storage and processing); B, chromatin structure and dynamics(information storage and processing); K, transcription (information storage and processing); H, coenzyme transport and metabolism (metabolism); R, general function prediction only (poorly characterized); F, nucleotide transport and metabolism (metabolism); S, function unknown (poorly characterized); L, replication, recombination, and repair (information storage and processing); E, amino acid transport and metabolism (metabolism); O, posttranslational modification, protein turnover, and chaperones (cellular processes and signaling); I, lipid transport and metabolism (metabolism); D, cell cycle control, cell division, and chromosome partitioning (cellular processes and signaling); J, translation, ribosomal structure, and biogenesis (information storage and processing); M, cell wall/membrane/envelope biogenesis (cellular processes and signaling); Q, secondary metabolite biosynthesis, transport, and catabolism (metabolism); T, signal transduction mechanisms (cellular processes and signaling); C, energy production and conversion (metabolism); V, defense mechanisms (cellular processes and signaling); G, carbohydrate transport and metabolism (metabolism); P, inorganic ion transport and metabolism (metabolism); Z, cytoskeleton (cellular processes and signaling); U, intracellular trafficking, secretion, and vesicular transport (cellular processes and signaling); N, cell motility (cellular processes and signaling).
As the transcription promoters well as the translation initiation signals for leaderless genes, the TA-like signals detected in bacteria merit a conscientious attention. In fact as shown in Figure
Atypical genes in prokaryotes
Besides the typical TA-like and the SD-like signals, the algorithm also identifies a lot of so-called atypical signals. Genes bearing these signals, namely atypical genes, are probably SD-less leadered genes, however they could also be leaderless genes with unknown promoter signals. In fact, several clades, for example
To sum up as a result, our analysis described above revealed that the translation initiation mechanism in bacteria is far from a simple scenario as previously imaged, and therefore a complex scenario with diversity should be rebuilt. Though SD-led genes are dominant in
Evolution scenario of translation initiation mechanisms
It is believed that the mechanism of translation initiation has dynamically changed during the prokaryotic genome evolution
Figure 5
Substitution rate of translation initiation signals
Substitution rate of translation initiation signals. The
The distance calculation gives a quantitative description for the substitution rate of translation initiation signals, but it does not give any evolutionary direction. Is there any trend in the macroevolution of translation initiation mechanism? To this end, a tree of the 16S rRNA sequences of 903 bacterial genomes was built and rooted with archaea outgroups. As shown in Figure
Figure 6
Trend in the evolution of leaderless genes
Trend in the evolution of leaderless genes. 903 bacterial genomes are put into 15 bins according to their distances to the root (between bacteria and archaea) on 16S rRNA tree. Average proportion of leaderless genes in each bin is calculated. The proportion of leaderless gene shows a rapid decrease followed by fluctuation at a low level.
There is a further question of concern, why leaderless genes were lost in many bacterial genomes while SD-led genes became dominant? One main driving force might be the coevolution of translation initiation signals and the operon structure
Figure 7
Coevolution of translation initiation signals and operons
Coevolution of translation initiation signals and operons. Each bacterial genome is denoted by two points in the figure (black and red), while each archaeal genome is denoted by a green point and a blue point. The proportion of operon distal genes is positively correlated to SD-led genes while negatively correlated to TA-led genes in both bacteria and archaea. Linear regressions are
One limitation of the current study should be noticed that only leaderless genes with transcriptional promoters resembling the Pribnow box are identified. This may cause an underestimation for the number of leaderless genes, especially in genomes with a large amount of atypical genes, such as
Although leaderless genes have been studied extensively in archaea, they have long been regarded as rare events in bacteria
In our method, all weight matrices and position distributions are learned automatically from the genome sequences. It's worth noting that there are several additional predefined parameters for the model, such as the length of region upstream to TIS in which signals are searched (denoted as
According to current evidences, SD-led initiation and leaderless initiation are probably both used by the LUCA. Evidence for the usage of SD-led initiation by the LUCA includes broad usage of SD sequence and the high conservation of anti-SD sequence in both bacteria and archaea
In this paper, we have studied the translation initiation signals in 1025 sequenced prokaryotic genomes and demonstrated the distribution of translation initiation mechanisms used in these genomes. The most surprising finding is that, though not as common as in archaea, there are substantial numbers of leaderless genes in bacteria. Most genomes with high percentage of leaderless genes are located near the root on the 16S rRNA tree and have relatively small number of operon distal genes. These facts show that current leaderless genes in bacteria are likely to be remnants of the ancestor and are retained because those genomes have low demand of organizing genes into operons for highly efficient and specialized gene expression.
953 bacterial and 72 archaeal genomes and their coding gene annotations in this study were downloaded from RefSeq in 2010
The multi-signal model and the signal detection algorithm
For the purpose of studying the translational initiation mechanism of prokaryotes, a straight way is to find the sequence patterns in the TIS upstream regions for each genome. With the sequence patterns, an algorithm can then build one or more positional weight matrices (PWMs) to describe the aligned positional frequency corresponding to potential signals. Such a relation has been used in many computational studies of translation initiation mechanism
To detect a signal associated with leaderless genes, the algorithm first selects the search region for each genome and the reference signal consensus. In archaea, leaderless genes can be detected by scanning the reference consensus of TA-rich transcription promoters at about 30 bp upstream to the TISs. Since the promoter mostly occurs at 30 bp upstream to the TSSs, thus for a leaderless gene the TIS and TSS should coincide
The multi-signal model of translation initiation signals
Usually the translation initiation signals are conserved in both content and position. For example, in
To describe multiple signals, herein we assume that there are
The likelihood function for the whole set
We then use the EM algorithm to obtain the maximum-likelihood estimation, details of which are described in Additional File
Additional file 2
Details of the EM algorithm to find signals. This additional file is to describe details of the EM algorithm used in finding translation initiation signals.
Click here for file
Classification of translation initiation signals
We then build the links between the four PWMs for each genome and the translation initiation way, which may be used to classify a PWM as which category of signals. A one-by-one manual examining would certainly carry this point. However it is impracticable for large-scale analysis over thousands of PWMs and more than thousand genomes in current study. Therefore, a classifier is designed to computationally calculate each PWM against those widely-known initiation signals as well as the "Atypical" signals
The first category of signal is the SD sequence widely used in prokaryotes. On 16S rRNA, the sequence UCACCUCCUU near 3'-end is conserved across species
Our analysis of the distribution of all distances shows that it displays a bimodal distribution separated at approximately 4.0. Then we classify all the PWMs with
The second category of signals to be examined is TA signals, which are known to usually appear upstream to the leaderless genes as our main concern in the current study. Since bacteria and archaea use different transcription signals, TA-signals have to be identified separately in two kingdoms. For bacteria, the Pribnow box weight matrix (6 × 4) is calculated by our algorithm from
All the distances also follow a bimodal distribution and the threshold is selected to be 0.8. Therefore the PWMs with TA_dis < 0.8 are regarded as TA-like signals.
In summary, for each PWM from a genome, it is first judged whether can be classified as SD-like signal, then TA-like signal. If neither can be decided, the PWM is regarded as the so-called atypical signal.
Classification of genes
Our basic working hypothesis for the model is that each sequence upstream to TIS of a gene has only one or none signal of translation initiation. So it is important to find out which signal each sequence has, this leads to that the gene downstream to the signal is regarded as a leaderless gene or not. As we described in the supporting text, when the model parameters converge,
meanwhile the probability that sequence
For each sequence
Strategy for validation of the signal detection algorithm
To test the statistical significance of the signals, we use simulated data that retain dinucleotide frequencies of the original sequences. For each 20 bp TIS upstream sequence in a given genome, a corresponding sequence with exactly the same dinucleotide frequency are generated using uShuffle
Estimation of TIS signal evolution distance
16S rRNA sequences were extracted from RefSeq annotation. After discarding those sequences that are longer than 1600 bp or are shorter than 1400 bp or have too much divergence with other sequences, 967 sequences were left, with 903 bacterial and 64 archaeal genomes. A list of the 967 genomes can be found at our webpage
For two genomes
(UTR): Un-translated region; (TIS): translation initiation site; (TSS): transcription start site; (SD): Shine-Dalgarno; (LUCA): Last Universal Common Ancestor; (COG): Clusters of Orthologous Groups.
Competing interests
The authors declare that they have no competing interests.
Authors' contributions
XBZ, GQH and HQZ conceived the study, XBZ and GQH designed the data analysis, XBZ and HQZ draft the manuscript, ZSS and HQZ cosupervised the progress of the work. All authors read and approved the final manuscript.
We thank Jiao Qu, Yongchu Liu, Hong Kang, Jiangtao Guo, Binbin Lai, Feifei He and others for beneficial discussions and helps to the work. We are grateful to Dr. Iain C Bruce of Zhejiang University School of Medicine for his linguistic help. The work was supported by the National Natural Science Foundation (30970667, 30770499 and 10721403) of China and the National Basic Research Program of China (2011CB707500).
Toward Automatic Reconstruction of a Highly Resolved Tree of Life
Dynamic evolution of translation initiation mechanisms in prokaryotes
The 3'-terminal sequence of Escherichia coli 16S ribosomal RNA: complementarity to nonsense triplets and ribosome binding sites
Autoregulation and function of a repressor in bacteriophage lambda
Leaderless mRNAs in bacteria: surprises in ribosomal recruitment and translational control
Evidence for the translation initiation of leaderless mRNAs by the intact 70 S ribosome without its dissociation into subunits in eubacteria
Selective stimulation of translation of leaderless mRNA by initiation factor 2: evolutionary implications for translation
A leaderless mRNA can bind to mammalian 80S ribosomes and direct polypeptide synthesis in the absence of translation initiation factors
Begin at the beginning: evolution of translational initiation
Two different and highly organized mechanisms of translation initiation in the archaeon Sulfolobus solfataricus
Leaderless transcripts of the crenarchaeal hyperthermophile Pyrobaculum aerophilum
Divergent transcriptional and translational signals in Archaea
Experimental characterization of Cis-acting elements important for translation and transcription in halophilic archaea
Correlations between Shine-Dalgarno sequences and gene features such as predicted expression levels and operon structures
Analysis of SD sequences in completed microbial genomes: Non-SD-led genes are as common as SD-led genes
The Translation initiation Signal Database
Anatomy of Escherichia coli sigma70 promoters
Translation of vph mRNA in Streptomyces lividans and Escherichia coli after removal of the 5' untranslated leader
A developmentally regulated gene encoding a repressor-like protein is essential for sporulation in Streptomyces coelicolor A3(2)
In vivo analysis of HPr reveals a fructose-specific phosphotransferase system that confers high-affinity uptake in Streptomyces coelicolor
Cascade of sigma factors in streptomycetes: identification of a new extracytoplasmic function sigma factor sigmaJ that is under the control of the stress-response sigma factor sigmaH in Streptomyces coelicolor A3(2)
Genomics of Actinobacteria: tracing the evolutionary history of an ancient phylum
Eubacterial, archaebacterial, and eukaryotic genes that encode leaderless mRNA
Evolution of translational initiation: new insights from the archaea
The COG database: an updated version includes eukaryotes
A novel mechanism for translation initiation operates in haloarchaea
Sequence features surrounding the translation initiation sites assigned on the genome sequence of Synechocystis sp. strain PCC6803 by amino-terminal protein sequencing
Regulation of translation via mRNA structure in prokaryotes and eukaryotes
Operon prediction using both genome-specific and general genomic information
A novel method for accurate operon predictions in all sequenced prokaryotes
The Complete Genome Sequence of the Lactic Acid Bacterium Lactococcus lactis ssp. lactis IL1403
An unexpected type of ribosomes induced by kasugamycin: a look into ancestral times of protein synthesis?
The sigma70 family of sigma factors
Repair of irradiation transforming deoxyribonucleic acid in wild type and a radiation-sensitive mutant of Micrococcus radiodurans
The genome sequence of the extreme thermophile Thermus thermophilus
Resistance of Lambda-Ci Translation to Antibiotics That Inhibit Translation Initiation
Determination of the optimal aligned spacing between the Shine-Dalgarno sequence and the translation initiation codon of Escherichia coli mRNAs
NCBI reference sequences (RefSeq): a curated non-redundant sequence database of genomes, transcripts and proteins
ProTISA: a comprehensive resource for translation initiation site annotation in prokaryotic genomes
Prediction of translation initiation site for microbial genomes with TriTISA
MED: a new non-supervised gene prediction algorithm for bacterial and archaeal genomes
Computational evaluation of TIS annotation for prokaryotic genomes
Accuracy improvement for identifying translation initiation sites in microbial genomes
GeneMarkS: a self-training method for prediction of gene starts in microbial genomes. Implications for finding sequence motifs in regulatory regions
Identifying bacterial genes and endosymbiont DNA with Glimmer
PromEC: An updated database of Escherichia coli mRNA promoters with experimentally identified transcriptional start sites
uShuffle: a useful tool for shuffling biological sequences while preserving the k-let counts
ClustalW and ClustalX version 2
MEGA: A biologist-centric software for evolutionary analysis of DNA and protein sequences
Reconstructing evolutionary trees from DNA and protein sequences: paralinear distances
WebLogo: a sequence logo generator
Genetic and transcriptional analysis of absA, an antibiotic gene cluster-linked two-component system that regulates multiple antibiotics in Streptomyces coelicolor
DevA, a GntR-like transcriptional regulator required for development in Streptomyces coelicolor
Beta-ketoacyl acyl carrier protein synthase III (FabH) is essential for fatty acid biosynthesis in Streptomyces coelicolor A3(2)
Regulation of the furA and catC operon, encoding a ferric uptake regulator homologue and catalase-peroxidase, respectively, in Streptomyces coelicolor A3(2)
Autophosphorylation of a bacterial serine/threonine kinase, AfsK, is inhibited by KbpA, an AfsK-binding protein
Substrate induction and glucose repression of maltose utilization by Streptomyces coelicolor A3(2) is controlled by malR, a member of the lacl-galR family of regulatory genes
Binding of PhoP to promoters of phosphate-regulated genes in Streptomyces coelicolor: identification of PHO boxes
Characterization of an inducible vancomycin resistance system in Streptomyces coelicolor reveals a novel gene (vanK) required for drug resistance


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Wei Jing Ge Map
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WOOSTER -- Bob Swartz and Cindy Farnsworth of Wooster announce the engagement of their daughter, Natalie Swartz, Wooster, to Michael Wrather of Wooster. He is the son of Dave and Jill Wrather, Smithville.
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What does it mean to be authentic? If we look it up in the dictionary, we'll find definitions such as these:
- "True to one's own personality, spirit, or character"
- "Conforming to reality and therefore worthy of trust, reliance, or belief"
- "Not false or copied"
This may be simple enough to understand from a theoretical standpoint, but how do we actually translate theory into practice when it comes to creating an authentic social media campaign?
The answers might surprise you. Be yourself. Embrace mistakes. Don't go for the best production value. Start with what you have. Be consistent with your message. Showcase your passion. Begin now.
In other words, authenticity is unpolished. Let me give you a quick example of what this all means.
Let's say you've been wanting to add video to your social media content strategy. However, you've been saying this for months now and you're waiting until the time is just right to start. Maybe you don't have the best camera. Maybe you want to wait until you can come up with better ideas for the video content. Maybe you don't have an ideal location to shoot in. Whatever the case may be, you're waiting for something.
This scenario happens all the time, especially with social media marketing campaigns. People want to wait until they feel like they've mastered everything or they need to wait until they have the latest and greatest technology and equipment.
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Long-term Evolution of the Outgassing of Comet Hale-Bopp From Radio Observations
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C/1995 O1 (Hale-Bopp) has been observed on a regular basis since August 1995 at millimetre and submillimetre wavelengths using IRAM, JCMT, CSO and SEST radio telescopes. The production rates of eight molecular species (CO, HCN, CH3OH, H2CO,H2S, CS, CH3CN,HNC) have been monitored as a function of heliocentric distance(rh from 7 AU pre-perihelion to 4 AU post-perihelion. As comet Hale-Bopp approached and receded from the Sun, these species displayed different behaviours. Far from the Sun, the most volatile species were found in general relatively more abundant in the coma. In comparison to other species, HNC, H2CO and CS showed a much steeper increase of the production rate with decreasing rh. Less than 1.5 AU from the Sun, the relative abundances were fairly stable and approached those found in other comets near 1 AU.
The kinetic temperature of the coma, estimated from the relative intensities of the CH3OH and CO lines, increased with decreasing rh, from about10 K at 7 AU to 110 K around perihelion. The expansion velocity of the gaseous species, derived from the line shapes, also increased with a law close torh3.
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