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Articles of Significant Interest Selected from This Issue by the Editors

While vaccinia virus (VACV) was used to eradicate smallpox worldwide, little is known about the origins of this virus or the many strains produced over more than a century of its propagation. Qin et al. (p. 1809 –1824) use shared genome features and idiosyncratic sequence differences to define the relatedness between available VACV strains. The data suggest that recombination and selection for essential genes near the telomeres of a horse pox-like ancestor combined with random sampling of nonclonal virus stocks likely drove VACV evolution and diversification.