タン, きも, い, まこと

gall bladder, courage, pluck, nerve

JLPT1 G7 S9 F1449

As per # (long and flat) + flesh → a long, flat piece of fat or grease → lip gloss (← lip gloss made from fat/grease). However, was adopted as a Shinjitai (new character form) to replace , according to which bears the meanings *gallbladder*; *courage*; *pluck* and *nerve*.

[Full Kanji Etymology List]