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Synopsis of the species of Myxobolus (Cnidaria, Myxozoa, Myxosporea) described between 2014 and 2020

A synopsis of the species of Myxobolus Bütschli, 1882 (Cnidaria, Myxosporea, Myxobolidae) described from 2014 up till now is presented. It includes 122 nominal species described all over the world. For each of the species, the most relevant morphological and morphometric data, as well as data are provided related to the location in the host, type host and type locality. The GenBank accession numbers are provided whenever possible, and the spores were redrawn based on the original descriptions. The bibliography includes all the papers containing the species descriptions.

The genus Myxobolus Bütschli, 1882 is between the most diversified Cnidaria of the class Myxozoa. They are mainly fish parasites and have a huge number of different host species, both freshwater and marine ones. Besides Myxobolus cerebralis (Hofer, 1903), causing the well-known whirling disease, some other species of Myxobolus have also proved to be pathogenic to their hosts Griffin 2010, Kaur et al. 2015). Eiras et al. (2005Eiras et al. ( , 2014 published two synopses of Myxobolus, which together attained 856 valid species. These papers proved to be useful for researchers interested in the group as judged by the high number of citations they reached -some 358 according to Research-Gate or 399 according to Google Scholar. Driven by the wide acceptance of this type of synopsis by the scientific community of the area, this paper has as aim to compilate the species of Myxobolus described from 2014 to date. The authors hope the present compilation is a useful tool for the researchers interested in this interesting group of parasites.

MATERIALS AND METHODS
The present list is based on an extensive internet search, as complete as possible, to detect all the species described since 2014. The authors recognise that, despite their efforts, some species were probably inadvertently overlooked, especially those in papers published in local journals, but hope that such omissions have been kept to a minimum.
The data are organised in tabulated format including the species name, authority, type host and type locality whenever possible, location in the host, description of vegetative stages and spores, including dimensions, and GenBank accession numbers when available.
Spores were re-drawn from the original illustrations, but some were not presented to scale as appeared in the original publications. A complete reference list including all the papers containing the species descriptions is presented. Fish names in Table 1 follow FishBase (Froese and Pauly 2019).
number of description of new species. In the first synopsis, there was a mean of 12.4 new species/year (112 species/ nine years), while now there is an increase to 20.5 species/ year (122 species/seven years). This increase in the pace of description of species suggests that there is still an undescribed diversity of species of Myxobolus to be revealed, and the success in this task will depend on the amount of researchers involved in this research area.
Concerning the 122 species presented in this synopsis, it is important to note that almost half (47%) were reported from two country -i.e., India with 35 species and Brazil with 23. This is probably due to two facts: (i) the diversity of fishes is very high in both countries, and (ii) there are several research groups well-established and involved in prospecting myxosporean diversity. Therefore, it cannot be concluded that Myxobolus spp. are more abundant (in relative terms considering the number of fish species within a country) if a similar research effort were done in some other countries with high fish diversity.