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Re-discovery of the type material of Leptoseris fragilis (Cnidaria, Anthozoa, Scleractinia) from the upper mesophotic in La Runion, a taxonomic puzzle.

Significant advances in the revision of the classification of scleractinian corals were made in the last years due to the use of an integrated systematics approach to resolve species boundaries and phylogenetic relationships (Kitahara et al. 2016). They are based on, yet not limited to, morphological and molecular analyses of specimens from multiple localities and the study of the type and historical specimens from museum collections (e.g. Benzoni et al. 2010, 2016). However, some coral lineages are still under study and remain unrevised taxonomically. The Agariciidae Gray, 1847 is one of those lineages. Including zooxanthellate and azooxanthellate taxa occurring from shallow to deep coral ecosystems worldwide (Veron 2000), Agariciidae are a typical component of shallow and Mesophotic Coral Ecosystems (MCEs) (Loya et al. 2019). In particular, agariciids of the genus Leptoseris Milne Edwards & Haime, 1849 are abundant, diverse and can dominate the benthic assemblages below 30 m depth in the Indo-Pacific, yet they are seldom identified to species level. In fact, their species boundaries are poorly understood and most observed morphologies in situ remain reported as Leptoseris spp. (e.g. Tamir et al. 2019, Hoarau et al. 2021). This uncertainty even remains when specimens are collected. It stems from a classification that is still largely based on morphological characters known to be uninformative and plagued by homoplasy (Kitahara et al. 2016). The few molecular phylogenies of Agariciidae published to date indicate that two of