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Patriaspirifer, a new genus of lower Devonian spiriferid brachiopods

IN 1944 G. A. Cooper assigned the well-known Oriskany species Spirifer murchisoni Castelnau to Acrospirifer Helmbrecht and Wedekind, 1923 (type species Spirifera primaeva Steininger, 1853). The generic name Acrospirifer subsequently became es­ tablished in eastern North America and later in Nevada (John­ son, 1970) where it was applied to species of an eastern North American enclave, which existed in the western United States for a time during the mid Early Devonian. In effect, Acrospirifer became a brachiopod cosmopolite. However, the literature, for a number of years, has contained information suggesting that the American and European species differ in some significant characteristics. The type species of Acrospirifer has an ornament of capillae and fila (Vandercammen, 1963, PI. 1, fig. 19, PI. 2, fig. 11); the American species are fimbriate. Although the sig­ nificance of this difference in ornament has been known since the time of Hall and Clarke (1893, p. 12-21), it did not signif­ icantly impact spiriferid systematics in the brachiopod Treatise (Pitrat, 1965). Revision of the Treatise now in progress will accord taxonomic significance to differences in shell fabric that result in capillate and fimbriate ornament. Thus, the fimbriate homeomorph of the capillate genus Acrospirifer is proposed as a new genus, Patriaspirifer.