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Primrose homostyles: A classic case of possible balancing selection revisited

In this issue of Molecular Ecology, Mora‐Carrera et al. (2022) revisit a case of the loss of an outcrossing system in primroses, which has been studied as an example of balancing selection in the wild since the 1940s. Molecular variants in the gene involved in the mutant self‐fertile phenotype, which is now known, help towards understanding this textbook example of breakdown of an outcrossing system. However, as often happens, new information also raises further questions.