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Reply [to “Comment on ‘Stochastic analysis of virus transport in aquifers,’ by Linda L. Campbell Rehmann, Claire Welty, and Ronald W. Harvey”]

suspended colloids or particles from lower-velocity regions of pore space due to the size of the suspended material particle." The phenomenon that Ginn defines as size exclusion, generally referred to as hydrodynamic hromatography [Small, 1974], is not referred to anywhere in our work. We point out that the term "size exclusion," as we have used the term, has been used extensively in the colloid transport literature to indicate as defined by Rehmann et al. [1999, p. 1987] that "colloids, unable to fit into the smaller pores, are limited to transport in larger ones" [e.g., Kretzschmar et al., 1997; McKay et al., 1993; Harvey, 1991, 1997]. Bales et al. [1989] have used the term "volume exclusion" to describe this phenomenon. The primary mechanisms that can result in virus breakthrough preceding that of a conservative tracer are: (1) what we define as [volume] size exclusion (exclusion of colloids from smaller pores due to their inability to fit into them); (2) preferential flow through highconductivity regions; and (3) what Ginn defines as size exclusion or hydrodynamic hromatography (exclusion of colloids from the lower-velocity regions of a pore throat due to their size). We agree with Ginn that these processes and the extent to which they affect colloid transport in porous media are poorly understood, and as stated previously, our work does not include the phenomenon defined by Ginn as size exclusion. It was our intention to investigate the extent to which the incorporation of heterogeneous aquifer parameters (i.e., hydraulic conductivity, pore water velocity, porosity, colloid filtration, etc.) affect virus transport behavior such as early virus breakthrough. Ginn further states that equation (la) of Rehmann et al. [1999] is free of a porosity transport operator. However, the development of equation (la) results in the presence of porosity in two key terms: first, the filtration term, and second, the detachment erm. Porosity has been perturbed in both of these parameter groupings in our analysis as well as in the local attachment equation (lb), both of which affect the resulting mean equation (30a) for free virus transport; we speculate that it is the perturbation of porosity in the filtration term that, in fact, can result in the early breakthrough behavior. Although