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Granule cells of the olfactory tubercle and the question of the islands of calleja

The granule cell clusters in the rat olfactory tubercle were studied in Nissl‐stained and Golgi‐impregnated sections. Discrete cell clusters that vary in size and shape occur mainly in the multiform layer and less often in the molecular layer. In cell‐stained sections they consist of small, round granule cells, 5–8 μm in diameter, that often surround a core or hilar area, which may contain larger neurons. In Golgi sections, the uni‐ or bipolar granule cells have a globular‐shaped soma and varicose dendrites that are thin, have few branches, and are usually less than 100 μm long. The dendrites remain within the border of the cluster. There are few spines on most granule cells; however, a small population of granule cells is spine‐rich. The axons are beaded, seldom have collaterals, and do not appear to exit from the cluster.