He earned a bachelor's degree with honors at the University of Washington in 1928 and a doctorate in physiology from Oxford University 1932 where he studied under Nobel Prize winner Sir Charles Sherrington.
He earned a bachelor's degree with honors at the University of Washington in 1928 from Oxford University 1932 where he studied under Nobel Prize winner Sir Charles Sherrington. He earned a bachelor's degree with honors at the University of a doctorate in physiology from Oxford University 1932 where he studied under Nobel Prize winner Sir Charles Sherrington.