Therefore, it is in the queen's interests to have her sons propagate the colony; by contrast, the worker will share half of her genes with her sons and only a quarter with her nearest brother.
It is in the queen's interests to have her sons propagate the colony. This is therefore. The worker will share half of her genes with her sons with her nearest brother. This is by contrast. The worker will share half of her genes with only a quarter with her nearest brother. This is by contrast.