From 1873 to 1886 he was an associate professor of anatomy and anthropology at Leipzig, and in 1886 he became a professor of anatomy and head of the anatomical institute at the University of Dorpat.
He was an associate professor of anatomy and anthropology at Leipzig. This was from 1873 to 1886. He became a professor of anatomy and head of the anatomical institute at the University of Dorpat. This was in 1886.