He graduated in 1869 at the Michigan Agricultural College, studied at Harvard under Prof. Asa Gray, in 1872 and in 1875-76, and was professor of botany at the Iowa Agricultural College from 1870 to 1884.
He graduated in 1869 at the Michigan Agricultural College, studied at Harvard under Prof. Asa Gray and in 1875-76. This was in 1872. He was professor of botany at the Iowa Agricultural College from 1870 to 1884.