After qualifying in medicine at University College Hospital Medical School, he returned to Cambridge for his PhD in immunology and was elected to a Fellowship at Trinity in 1973 for his discovery of the role of complement in induction of antibody production.
He returned to Cambridge for his PhD in immunology. He was elected to a Fellowship at Trinity in 1973 for his discovery of the role of complement in induction of antibody production. He was qualifying in medicine at University College Hospital Medical School.