From 1889 to 1891 he studied medicine at University College, Liverpool, where he was appointed in 1891 Holt Fellow in Physiology and did research under Francis Gotch from 1891 to 1897.
From 1889 to 1891 he studied medicine at University College, Liverpool. In 1891, he was appointed Holt Fellow in Physiology at University College, Liverpool, and did research under Francis Gotch from 1891 to 1897.