Instruct: Given a question, retrieve documents that can help answer the question.
Query: Where were fossils of the Smilodon first discovered?
During the 1830s, Danish naturalist Peter Wilhelm Lund and his assistants collected fossils in the calcareous caves near the small town of Lagoa Santa, Minas Gerais, Brazil. Among the thousands of fossils found, he recognized a few isolated cheek teeth as belonging to a hyena, which he named "Hyaena neogaea" in 1839. After more material was found (including canine teeth and foot bones), Lund concluded the fossils instead belonged to a distinct genus of felid, though transitional to the hyenas. He stated it would have matched the largest modern predators in size, and was more robust than any modern cat. Lund originally wanted to name the new genus "Hyaenodon", but realizing this had recently become preoccupied by another prehistoric predator, he instead named it "Smilodon populator" in 1842. He explained the Ancient Greek meaning of "Smilodon" as ("smilē"), a scalpel or two-edged knife, and "οδόντος" ("odontús"), tooth. This has also been translated as "tooth shaped like double-edged knife". He explained the species name "populator" as "the destroyer", which has also been translated as "he who brings devastation". By 1846, Lund had acquired nearly every part of the skeleton (from different individuals), and more specimens were found in neighboring countries by other collectors in the following years. Though some later authors used Lund's original species name "neogaea" instead of "populator", it is now considered an invalid "nomen nudum" ("naked name"), as it was not accompanied with a proper description and no type specimens were designated. Some South American specimens have been referred to other genera, subgenera, species, and subspecies, such as "Smilodontidion riggii", "Smilodon" ("Prosmilodon") "ensenadensis", and "S. bonaeriensis", but these are now thought to be junior synonyms of "S. populator".
Whether "Smilodon" was sexually dimorphic has implications for its reproductive behavior. Based on their conclusions that "Smilodon fatalis" had no sexual dimorphism, Van Valkenburgh and Sacco suggested in 2002 that, if the cats were social, they would likely have lived in monogamous pairs (along with offspring) with no intense competition among males for females. Likewise, Meachen-Samuels and Binder (2010) concluded that aggression between males was less pronounced in "S. fatalis" than in the American lion. Christiansen and Harris (2012) found that, as "S. fatalis" did exhibit some sexual dimorphism, there would have been evolutionary selection for competition between males. The structure of the hyoid bones suggest that "Smilodon" communicated by roaring, like modern big cats. The ability to roar may have implications for the social life of "Smilodon".
The jaws of machairodonts, especially more derived species with longer canines, such as "Smilodon" and "Megantereon", are unusually weak. Digital reconstructions of the skulls of lions and of "Smilodon" show that the latter would have fared poorly with the stresses of holding onto struggling prey. The main issue was the stresses suffered by the mandible: a strong force threatened to break the jaw as pressure was placed on its weakest points.
"Sinosauropteryx" lived in what is now northeastern China during the early Cretaceous period. It was among the first dinosaurs discovered from the Yixian Formation in Liaoning Province, and was a member of the Jehol Biota. Well-preserved fossils of this species illustrate many aspects of its biology, such as its diet and reproduction.
The first publicized discovery of a partial fossil mosasaur skull in 1764 by quarry workers in a subterranean gallery of a limestone quarry in Mount Saint Peter, near the Dutch city of Maastricht, preceded any major dinosaur fossil discoveries, but remained little known. However, a second find of a partial skull drew the Age of Enlightenment's attention to the existence of fossilized animals that were different from any known living creatures. When the specimen was discovered between 1770 and 1774, Johann Leonard Hoffmann, a surgeon and fossil collector, corresponded about it with the most influential scientists of his day, making the fossil famous. The original owner, though, was Godding, a canon of Maastricht cathedral.
During the upper Triassic period, about 210 million years ago, the region around Frick was a dry lowland with flat hills and small depressions. During the rainy season, the depressions filled with water and dinosaurs congregated around the ponds. When they died, their bodies were covered by the mud in the ponds and fossilized, creating rich fossil beds in Frick. The first Plateosaurus fossils were discovered in 1961 and further excavations during the following decades have discovered numerous fossils. In 2006, the only Coelophysoidea (a small flesh-eating dinosaur species) fossil in Switzerland, was found by an amateur paleontologist in Frick.
Features necessary for defending a territory, such as the horns and broad-based skull of the modern black wildebeest, have been found in their fossil ancestors. The earliest known fossil remains are in sedimentary rock in Cornelia in the Orange Free State and date back about 800,000 years. Fossils have also been reported from the Vaal River deposits, though whether or not they are as ancient as those found in Cornelia is unclear. Horns of the black wildebeest have been found in sand dunes near Hermanus in South Africa. This is far beyond the recorded range of the species and it has been suggested that these animals may have migrated to that region from the Karoo.
Three specimens have been assigned to "Sinosauropteryx prima": the holotype GMV 2123 (and its counter slab [opposite face], NIGP 127586), NIGP 127587, and D 2141. Another specimen, IVPP V14202, was assigned to the genus but not to the only species by Zhang and colleagues. The assignment of an additional larger specimen to "S. prima", GMV 2124, was later found to be in error. All of the fossils were found in the Jianshangou or Dawangzhangzi Beds of the Yixian Formation in the Beipiao and Lingyuan regions of Liaoning, China. These fossil beds have been dated to 124.6–122 million years ago, during the early Aptian stage of the Early Cretaceous.
The team discovered the remains of the largest dinosaur to walk in the southern parts of Africa , namely a 190-million-year-old fossilised egg of a Massospondylus dinosaur. This dinosaur lived in the Late Triassic and Early Jurassic, a period from about 230-million years ago to about 185-million years ago. The Massospondylus wandered about in great herds, migrating back and forth between what would millions of years later become Southern Africa and Russia, when the southern super-continent, Gondwana, was still intact. The rocks that were laid down during this period are called the Stormberg Group of rocks, and it is in rocks of the Stormberg Group that the fossils of Massospondylus and other dinosaurs were found. Massospondylus dinosaurs hatched from eggs not much larger than a hen’s egg, but grew into big creatures 5 to 6 metres long. They had large bodies, long necks and small heads, and long tails.
Many important fossils have been found in the sandstones and clays of the Weald, including, for example, "Baryonyx". The famous scientific hoax of Piltdown Man was claimed to have come from a gravel pit at Piltdown near Uckfield. The first "Iguanodon" was identified after a Mary Mantell unearthed some fossilised teeth by a road in Sussex in 1822. Her husband, Gideon Mantell, noticed they were similar to modern iguana teeth but many times larger; this important find led to the discovery of dinosaurs.
220,000,000 BC, Late Triassic (Arizona) − This episode follows the evolution of the first dinosaurs and other prehistoric creatures in the late Triassic period.152,000,000 BC, Late Jurassic (Colorado) − This episode follows the story of a young female "Diplodocus" and her brothers and sisters in the late Jurassic period.149,000,000 BC, Late Jurassic (Oxfordshire) − This episode follows the life of several baby "Ophthalmosaurus" and a bull "Liopleurodon" in the late Jurassic period.127,000,000 BC, Early Cretaceous (Brazil) − This episode follows a male "Ornithocheirus" on his long journey to the mating grounds in the early Cretaceous period.106,000,000 BC, Early Cretaceous (Antarctica) − This episode follows the hardships of an Antarctican "Leaellynasaura" clan in the early Cretaceous period.65,500,000 BC, Late Cretaceous (Montana) − This episode follows the final years of the dinosaurs and a mother "Tyrannosaurus rex" and her babies in the late Cretaceous period.49,000,000 BC, Early Eocene (Germany) − This episode follows the life of a mother "Leptictidium" and her babies in the dense forests of Germany in the early Eocene epoch.36,000,000 BC, Late Eocene (Pakistan) − This episode follows the sea journey of a pregnant female "Basilosaurus" search for food in the late Eocene epoch.25,000,000 BC, Late Oligocene (Mongolia) − This episode follows the life of a young "Paraceratherium" and his mother from his birth in the late Oligocene epoch.3,200,000 BC, Late Pliocene (Ethiopia) − This episode follows the hardships of an "Australopithecus" group and a young called Blue in the late Pliocene epoch.1,000,000 BC, Early Pleistocene (Paraguay) − This episode follows a male "Smilodon" called Half-Tooth after he lost his territory in the early Pleistocene epoch.30,000 BC, Late Pleistocene (North Sea, Belgium, and Swiss Alps) − This episode follows a herd of woolly mammoth journey to the Alps in the last Ice Age of the late Pleistocene epoch.This episode details a list of animals from the first season that the developers thought were the most impressive creatures on Earth.
Recent molecular phylogenetic studies suggest that most placental orders diverged late in the Cretaceous period, about 100 to 85 million years ago, but that modern families first appeared later, in the late Eocene and early Miocene epochs of the Cenozoic period. Fossil-based analyses, on the contrary, limit the placentals to the Cenozoic. Many Cretaceous fossil sites contain well-preserved lizards, salamanders, birds, and mammals, but not the modern forms of mammals. It is likely that they simply did not exist, and that the molecular clock runs fast during major evolutionary radiations. On the other hand, there is fossil evidence from of hoofed mammals that may be ancestors of modern ungulates.
In 1983, a relatively complete skeleton was excavated from the Smokejacks pit in Surrey, Engand. These remains were described by British paleontologists Alan J. Charig and Angela C. Milner in 1986 as the holotype of a new species, "Baryonyx walkeri." After the discovery of "Baryonyx," many new genera have since been described, the majority on very incomplete remains. However, other finds bear enough fossil material and distinct anatomical features to be assigned with confidence. Paul Sereno and colleagues described "Suchomimus" in 1998"," a baryonychine from Niger on the basis of a partial skeleton found in 1997. Later in 2004, partial jaw bones were recovered from the Alcântara Formation, these were referred to a new genus of spinosaurine named "Oxalaia" in 2011 by Alexander Kellner.
"Pteranodon" was the first pterosaur found outside of Europe. Its fossils first were found by Othniel Charles Marsh in 1870, in the Late Cretaceous Smoky Hill Chalk deposits of western Kansas. These chalk beds were deposited at the bottom of what was once the Western Interior Seaway, a large shallow sea over what now is the midsection of the North American continent. These first specimens, YPM 1160 and YPM 1161, consisted of partial wing bones, as well as a tooth from the prehistoric fish "Xiphactinus", which Marsh mistakenly believed to belong to this new pterosaur (all known pterosaurs up to that point had teeth). In 1871, Marsh named the find ""Pterodactylus oweni"", assigning it to the well-known (but much smaller) European genus "Pterodactylus". Marsh also collected more wing bones of the large pterosaur in 1871. Realizing that the name he had chosen had already been used for Harry Seeley's European pterosaur species "Pterodactylus oweni" in 1864, Marsh re-named his giant North American pterosaur Pterodactylus occidentalis, meaning "Western wing finger," in his 1872 description of the new specimen. He also named two additional species, based on size differences: Pterodactylus ingens (the largest specimen so far), and Pterodactylus velox (the smallest).
The first documented fossils to be recovered from the Bridger Formation were discovered on 11 August 1849 by Captain Howard Stansbury, who documented the discovery of fossilized shells and wood in his expedition report while scouting out the region for the United States Army Corps of Topographical Engineers. In the early-1860s, trapper Jack Robinson claimed to have discovered a number of sites along the base of the Uinta Mountains where grizzly bears had been turned to stone. When these claims were called into question by judge William A. Carter, Robinson brought Carter a bag filled with the fossils. One of the specimens recovered by Robinson was a well-preserved skull which resembled that of a grizzly bear. Judger Carter invited Louis Agassiz to observe the local strata, but Agassiz declined as the journey would have involved riding horseback to the site, a mode of transportation Agassiz abhorred. Carter's son-in-law, Dr. J. Van A. Carter, would go on to send a number of fossils to palaeontologist Joseph Leidy at the University of Pennsylvania in 1869. These fossils included the first Bridgerian fossil taxa, "Omomys carteri"; and the skull discovered by Robinson, which was described as "Palaeosyops paludosus". Another researcher responsible for sending off specimens was Dr. Joseph K. Corson, a close friend of Leidy's who hosted him and his family on two three to Fort Bridger in 1872, 1873, and 1879.
The museum's five rooms shows paleontological finds from the Besano area, including the "Besanosaurus", a type of ichthyosaur from the middle Triassic period.
The 6 meter "Besanosaurus leptorhynchus" discovered in 1993 is the biggest sea reptile fossil found in Italy, and was probably a pregnant female.
Since 2001 the museum has also housed the remains of a "Saltriosaurus", discovered in a quarry in Saltrio, which is the first big carnivorous dinosaur found in Lombardia.
A number of partial fossil remains of "Secodontosaurus" have been identified from its characteristic long skull and jaws. The postcranial skeletal material from different individuals includes parts of the backbone with clear evidence of a tall sail very similar to that of "Dimetrodon". The limbs and tail are incomplete but probably resembled those of "Dimetrodon" as well. Like "Dimetrodon", "Secodontosaurus" would have had a short neck, robust body, short limbs, and a long tail. In light of such similarities, some skeletal remains with missing or fragmentary skulls that were previously identified as "Dimetrodon" may in fact belong to "Secodontosaurus". A key noncranial difference can be found in the axis neck vertebra, which has a tall and broad neural spine in "Dimetrodon" but has a lower neural spine in "Secodontosaurus".
The tail of "Dimetrodon" makes up a large portion of its total body length and includes around 50 caudal vertebrae. Tails were missing or incomplete in the first described skeletons of "Dimetrodon"; the only caudal vertebrae known were the eleven closest to the hip. Since these first few caudal vertebrae narrow rapidly as they progress farther from the hip, many paleontologists in the late nineteenth and early twentieth centuries thought that "Dimetrodon" had a very short tail. It was not until 1927 that a largely complete tail of "Dimetrodon" was described.
Paleontology has a long history in the State of Illinois, stretching at least as far back as the 1850s, when the first Mazon Creek fossils were being found. Early in the ensuing 20th century a Silurian age fossil reef system was discovered in the state. In the 1950s Francis Tully discovered the "monster" that would later be named in his honor. The Pennsylvanian species "Tullimonstrum gregarium" ("Tully Monster") is the Illinois state fossil. It is one of the few officially designated state fossils that is endemic to the state it represents.
The first "Hylaeosaurus" fossils were discovered in West Sussex. On 20 July 1832, fossil collector Gideon Mantell wrote to Professor Benjamin Silliman that when a gunpowder explosion had demolished a quarry rock face in Tilgate Forest, several of the boulders freed showed the bones of a saurian. A local fossil dealer had assembled the about fifty pieces, described by him as a "great consarn of bites and boanes". Having doubts about the value of the fragments, Mantell had nevertheless purchased the pieces and soon discovered they could be united into a single skeleton, partially articulated. Mantell was delighted with the find because previous specimens of "Megalosaurus" and "Iguanodon" had consisted of single bone elements. The discovery in fact represented the most complete non-avian dinosaur skeleton known at the time. He was strongly inclined to describe the find as belonging to the latter genus, but during a visit by William Clift, the curator of the Royal College of Surgeons of England museum, and his assistant John Edward Gray, he began to doubt the identification. Clift was the first to point out that several plates and spikes were probably part of a body armour, attached to the back or sides of the rump. In November 1832 Mantell decided to create a new generic name: "Hylaeosaurus". It is derived from the Greek ὑλαῖος, "hylaios", "of the wood". Mantell originally claimed the name "Hylaeosaurus" meant "forest lizard", after the Tilgate Forest in which it was discovered. Later, he claimed that it meant "Wealden lizard" ("wealden" being another word for "forest"), in reference to the Wealden Group, the name for the early Cretaceous geological formation in which the dinosaur was first found.
This species of wildebeest seems to have evolved around 2.5 million years ago. The black wildebeest is believed to have diverged from the blue wildebeest to become a distinct species around a million years ago, in the mid- to late Pleistocene. Fossil evidence suggests that blue wildebeest were quite common in the Cradle of Humankind in the past. Apart from eastern Africa, fossils are commonly found in Elandsfontein, Cornelia, and Florisbad.
The lineage of "Homotherium" is estimated (based on mitochondrial DNA sequences) to have diverged from that of "Smilodon" about 18 million years ago. "Homotherium" probably derived from "Machairodus" and appeared for the first time at the Miocene–Pliocene border, about 4 to 5 million years ago. During the Pleistocene it occurred in vast parts of Eurasia, North America and until the middle Pleistocene (about 1.5 million years ago) even in Africa. A fossil of "H. crenatidens" was inadvertently dredged from the bed of the North Sea, which was a flat, low-lying extent of marshy tundra laced with rivers during the last glacial period. There has also been a discovery of 1.8-million-year-old fossils in Venezuela, indicating that scimitar-toothed cats were able to invade South America along with "Smilodon" during the Great American Interchange. These remains form the holotype of "Homotherium venezuelensis". How long they lasted in South America is not yet evident. "Homotherium" survived in Eurasia until about 28,000 years ago.
Radiocarbon analysis of human remains from the cave returned dates of 2390+/-145 B.P. and 1690+/-115 B.P. These remains were collected from an upper zone approximately 16-feet above the saber-tooth bones. The human remains are believed to be from the Woodland Period and originated thousands of years after the "Smilodon" find. According to Guilday (1977), collagen from the "Smilodon" remains returned radiocarbon dates of 9410+/-155 B.P. and 10,035+/-650 B.P. These dates are extremely late for the presence of "Smilodon" in the Southeast, both contemporaneous with the Dalton horizon and overlapping Paleoindian occupations along the Cumberland River by at least 1,000 years. As such, the dates for the "Smilodon" remains from the First American site should be regarded with some skepticism.
"Smilodon" lived during the Pleistocene epoch (2.5 mya–10,000 years ago), and was perhaps the most recent of the saber-toothed cats. "Smilodon" probably lived in a closed habitat such as forest or bush. Fossils of the genus have been found throughout the Americas. In North America, the varied habitat supported other saber-toothed cats in addition to "Smilodon", such as "Homotherium" and "Xenosmilus"; the habitat here varied from subtropical forests and savannah in the south, to treeless mammoth steppes in the north. "Smilodon" inhabited the temperate latitudes of North America, where the mosaic vegetation of woods, shrubs, and grasses in the southwest supported large herbivores such as horses, bison, antelope, deer, camels, mammoths, mastodons, and ground sloths. Other large carnivores included dire wolves, short-faced bear ("Arctodus simus") and the American lion. Due to competition from larger carnivores in North America, "S. fatalis" was perhaps not able to attain the same size as "S. populator". The similar sizes of "S. fatalis" and the American lion suggests niche overlap and direct competition between these species, and they appear to have fed on similarly sized prey.
Several "Smilodon" fossils show signs of ankylosing spondylitis, hyperostosis and trauma; some also had arthritis, which gave them fused vertebrae. One study of 1,000 "Smilodon" skulls found that 30% of them had eroded parietal bones, which is where the largest jaw muscles attach. They also showed signs of microfractures, and the weakening and thinning of bones possibly caused by mechanical stress from the constant need to make stabbing motions with the canines. Bony growths where the deltoid muscle inserted in the humerus is a common pathology for a La Brea specimen, which was probably due to repeated strain when "Smilodon" attempted to pull down prey with its forelimbs. Sternum injuries are also common, probably due to collision with prey. Some bones also show evidence of having been bitten by other "Smilodon", with one skull showing an unhealed wound from a canine, seemingly fatal, and a scapula from the same site that had been pierced by a canine. The frequency of trauma in "S. fatalis" specimens was 4.3%, compared to 2.8% in the dire wolf, which implies the ambush predatory behavior of the former led to greater risk of injury than the pursuit predatory behavior of the latter. "Smilodon" remains exhibit relatively more shoulder and lumbar vertebrae injuries.