Instruct: Given a question, retrieve relevant documents that can help answer the question.
Query: When was the first Pteranodon fossil found?
"Pteranodon" was the first pterosaur found outside of Europe. Its fossils first were found by Othniel Charles Marsh in 1870, in the Late Cretaceous Smoky Hill Chalk deposits of western Kansas. These chalk beds were deposited at the bottom of what was once the Western Interior Seaway, a large shallow sea over what now is the midsection of the North American continent. These first specimens, YPM 1160 and YPM 1161, consisted of partial wing bones, as well as a tooth from the prehistoric fish "Xiphactinus", which Marsh mistakenly believed to belong to this new pterosaur (all known pterosaurs up to that point had teeth). In 1871, Marsh named the find ""Pterodactylus oweni"", assigning it to the well-known (but much smaller) European genus "Pterodactylus". Marsh also collected more wing bones of the large pterosaur in 1871. Realizing that the name he had chosen had already been used for Harry Seeley's European pterosaur species "Pterodactylus oweni" in 1864, Marsh re-named his giant North American pterosaur Pterodactylus occidentalis, meaning "Western wing finger," in his 1872 description of the new specimen. He also named two additional species, based on size differences: Pterodactylus ingens (the largest specimen so far), and Pterodactylus velox (the smallest).
It is based on the type specimen or holotype BPV-078, an incomplete skeleton and skull. Its skull, including a large crest, was toothless and similar to that of "Pteranodon". The skull lacks the point of the snout but it was in life less than 200 millimeters long (7.9 inches), and the animal had a wingspan of about 1.1 meters (3.6 feet). A second specimen, D2526, described in 2006, had a larger wingspan. Despite its similarities to "Pteranodon", "Eopteranodon" was not placed into a family by its describers, who put it into the clade Pteranodontia as "incertae sedis" (uncertain position).
"Sinosauropteryx" lived in what is now northeastern China during the early Cretaceous period. It was among the first dinosaurs discovered from the Yixian Formation in Liaoning Province, and was a member of the Jehol Biota. Well-preserved fossils of this species illustrate many aspects of its biology, such as its diet and reproduction.
Many important fossils have been found in the sandstones and clays of the Weald, including, for example, "Baryonyx". The famous scientific hoax of Piltdown Man was claimed to have come from a gravel pit at Piltdown near Uckfield. The first "Iguanodon" was identified after a Mary Mantell unearthed some fossilised teeth by a road in Sussex in 1822. Her husband, Gideon Mantell, noticed they were similar to modern iguana teeth but many times larger; this important find led to the discovery of dinosaurs.
220,000,000 BC, Late Triassic (Arizona) − This episode follows the evolution of the first dinosaurs and other prehistoric creatures in the late Triassic period.152,000,000 BC, Late Jurassic (Colorado) − This episode follows the story of a young female "Diplodocus" and her brothers and sisters in the late Jurassic period.149,000,000 BC, Late Jurassic (Oxfordshire) − This episode follows the life of several baby "Ophthalmosaurus" and a bull "Liopleurodon" in the late Jurassic period.127,000,000 BC, Early Cretaceous (Brazil) − This episode follows a male "Ornithocheirus" on his long journey to the mating grounds in the early Cretaceous period.106,000,000 BC, Early Cretaceous (Antarctica) − This episode follows the hardships of an Antarctican "Leaellynasaura" clan in the early Cretaceous period.65,500,000 BC, Late Cretaceous (Montana) − This episode follows the final years of the dinosaurs and a mother "Tyrannosaurus rex" and her babies in the late Cretaceous period.49,000,000 BC, Early Eocene (Germany) − This episode follows the life of a mother "Leptictidium" and her babies in the dense forests of Germany in the early Eocene epoch.36,000,000 BC, Late Eocene (Pakistan) − This episode follows the sea journey of a pregnant female "Basilosaurus" search for food in the late Eocene epoch.25,000,000 BC, Late Oligocene (Mongolia) − This episode follows the life of a young "Paraceratherium" and his mother from his birth in the late Oligocene epoch.3,200,000 BC, Late Pliocene (Ethiopia) − This episode follows the hardships of an "Australopithecus" group and a young called Blue in the late Pliocene epoch.1,000,000 BC, Early Pleistocene (Paraguay) − This episode follows a male "Smilodon" called Half-Tooth after he lost his territory in the early Pleistocene epoch.30,000 BC, Late Pleistocene (North Sea, Belgium, and Swiss Alps) − This episode follows a herd of woolly mammoth journey to the Alps in the last Ice Age of the late Pleistocene epoch.This episode details a list of animals from the first season that the developers thought were the most impressive creatures on Earth.
During the upper Triassic period, about 210 million years ago, the region around Frick was a dry lowland with flat hills and small depressions. During the rainy season, the depressions filled with water and dinosaurs congregated around the ponds. When they died, their bodies were covered by the mud in the ponds and fossilized, creating rich fossil beds in Frick. The first Plateosaurus fossils were discovered in 1961 and further excavations during the following decades have discovered numerous fossils. In 2006, the only Coelophysoidea (a small flesh-eating dinosaur species) fossil in Switzerland, was found by an amateur paleontologist in Frick.
The fact that females appear to have outnumbered males two to one suggests that, as with modern animals with size-related sexual dimorphism, such as sea lions and other pinnipeds, "Pteranodon" might have been polygynous, with a few males competing for association with groups consisting of large numbers of females. Similar to modern pinnipeds, "Pteranodon" may have competed to establish territory on rocky, offshore rookeries, with the largest, and largest-crested, males gaining the most territory and having more success mating with females. The crests of male "Pteranodon" would not have been used in competition, but rather as "visual dominance-rank symbols", with display rituals taking the place of physical competition with other males. If this hypothesis is correct, it also is likely that male "Pteranodon" played little to no part in rearing the young; such a behavior is not found in the males of modern polygynous animals who father many offspring at the same time.
The first publicized discovery of a partial fossil mosasaur skull in 1764 by quarry workers in a subterranean gallery of a limestone quarry in Mount Saint Peter, near the Dutch city of Maastricht, preceded any major dinosaur fossil discoveries, but remained little known. However, a second find of a partial skull drew the Age of Enlightenment's attention to the existence of fossilized animals that were different from any known living creatures. When the specimen was discovered between 1770 and 1774, Johann Leonard Hoffmann, a surgeon and fossil collector, corresponded about it with the most influential scientists of his day, making the fossil famous. The original owner, though, was Godding, a canon of Maastricht cathedral.
"Dimetrodon" was a quadrupedal, sail-backed synapsid. Most "Dimetrodon" species ranged in length from and are estimated to have weighed between . The largest known species of "Dimetrodon" is "D. angelensis" at and the smallest is "D. teutonis" at . The larger species of "Dimetrodon" were among the largest predators of the Early Permian, although the closely related "Tappenosaurus", known from skeletal fragments in slightly younger rocks, may have been even larger at an estimated in total body length. Although some "Dimetrodon" species could grow very large, many juvenile specimens are known.
The team discovered the remains of the largest dinosaur to walk in the southern parts of Africa , namely a 190-million-year-old fossilised egg of a Massospondylus dinosaur. This dinosaur lived in the Late Triassic and Early Jurassic, a period from about 230-million years ago to about 185-million years ago. The Massospondylus wandered about in great herds, migrating back and forth between what would millions of years later become Southern Africa and Russia, when the southern super-continent, Gondwana, was still intact. The rocks that were laid down during this period are called the Stormberg Group of rocks, and it is in rocks of the Stormberg Group that the fossils of Massospondylus and other dinosaurs were found. Massospondylus dinosaurs hatched from eggs not much larger than a hen’s egg, but grew into big creatures 5 to 6 metres long. They had large bodies, long necks and small heads, and long tails.
The Early Cretaceous Epoch spans from 145 million to 100 million years ago. The Early Cretaceous saw the expansion of seaways, and as a result, the decline and extinction of sauropods (except in South America). Many coastal shallows were created, and that caused ichthyosaurs to die out. Mosasaurs evolved to replace them as apex species of the seas. Some island-hopping dinosaurs, like "Eustreptospondylus", evolved to cope with the coastal shallows and small islands of ancient Europe. Other dinosaurs, such as "Carcharodontosaurus" and "Spinosaurus", rose to fill the empty space that the Jurassic-Cretaceous extinction had created. Of the most successful would be the "Iguanodon" which spread to every continent. Seasons came back into effect and the poles grew seasonally colder. Dinosaurs such as the "Leaellynasaura" inhabited the polar forests year-round, while many dinosaurs, such as the "Muttaburrasaurus", migrated there during summer . Since it was too cold for crocodiles, it was the last stronghold for large amphibians, such as the "Koolasuchus". In this epoch Pterosaurs reached their maximum diversity and grew larger, as species like "Tapejara" and "Ornithocheirus" took to the skies. The first true birds evolved, possibly sparking competition between them and the pterosaurs.
Baryonyx () is a genus of theropod dinosaur which lived in the Barremian stage of the Early Cretaceous Period, about 130–125 million years ago. The first skeleton was discovered in 1983 in the Weald Clay Formation of Surrey, England, and became the holotype specimen of B. walkeri, named by palaeontologists Alan J. Charig and Angela C. Milner in 1986. The generic name, "Baryonyx", means "heavy claw" and alludes to the animal's very large claw on the first finger; the specific name, "walkeri", refers to its discoverer, amateur fossil collector William J. Walker. The holotype specimen is one of the most complete theropod skeletons from the UK (and remains the most complete spinosaurid), and its discovery attracted media attention. Specimens later discovered in other parts of the United Kingdom and Iberia have also been assigned to the genus.
The wing shape of "Pteranodon" suggests that it would have flown rather like a modern-day albatross. This is based on the fact that "Pteranodon" had a high aspect ratio (wingspan to chord length) similar to that of the albatross — 9:1 for "Pteranodon", compared to 8:1 for an albatross. Albatrosses spend long stretches of time at sea fishing, and use a flight pattern called "dynamic soaring" which exploits the vertical gradient of wind speed near the ocean surface to travel long distances without flapping, and without the aid of thermals (which do not occur over the open ocean the same way they do over land). While most of a "Pteranodon" flight would have depended on soaring, like long-winged seabirds, it probably required an occasional active, rapid burst of flapping, and studies of "Pteranodon" wing loading (the strength of the wings vs. the weight of the body) indicate that they were capable of substantial flapping flight, contrary to some earlier suggestions that they were so big they could only glide.
Recent molecular phylogenetic studies suggest that most placental orders diverged late in the Cretaceous period, about 100 to 85 million years ago, but that modern families first appeared later, in the late Eocene and early Miocene epochs of the Cenozoic period. Fossil-based analyses, on the contrary, limit the placentals to the Cenozoic. Many Cretaceous fossil sites contain well-preserved lizards, salamanders, birds, and mammals, but not the modern forms of mammals. It is likely that they simply did not exist, and that the molecular clock runs fast during major evolutionary radiations. On the other hand, there is fossil evidence from of hoofed mammals that may be ancestors of modern ungulates.
The first documented fossils to be recovered from the Bridger Formation were discovered on 11 August 1849 by Captain Howard Stansbury, who documented the discovery of fossilized shells and wood in his expedition report while scouting out the region for the United States Army Corps of Topographical Engineers. In the early-1860s, trapper Jack Robinson claimed to have discovered a number of sites along the base of the Uinta Mountains where grizzly bears had been turned to stone. When these claims were called into question by judge William A. Carter, Robinson brought Carter a bag filled with the fossils. One of the specimens recovered by Robinson was a well-preserved skull which resembled that of a grizzly bear. Judger Carter invited Louis Agassiz to observe the local strata, but Agassiz declined as the journey would have involved riding horseback to the site, a mode of transportation Agassiz abhorred. Carter's son-in-law, Dr. J. Van A. Carter, would go on to send a number of fossils to palaeontologist Joseph Leidy at the University of Pennsylvania in 1869. These fossils included the first Bridgerian fossil taxa, "Omomys carteri"; and the skull discovered by Robinson, which was described as "Palaeosyops paludosus". Another researcher responsible for sending off specimens was Dr. Joseph K. Corson, a close friend of Leidy's who hosted him and his family on two three to Fort Bridger in 1872, 1873, and 1879.
The museum's five rooms shows paleontological finds from the Besano area, including the "Besanosaurus", a type of ichthyosaur from the middle Triassic period.
The 6 meter "Besanosaurus leptorhynchus" discovered in 1993 is the biggest sea reptile fossil found in Italy, and was probably a pregnant female.
Since 2001 the museum has also housed the remains of a "Saltriosaurus", discovered in a quarry in Saltrio, which is the first big carnivorous dinosaur found in Lombardia.
The first "Hylaeosaurus" fossils were discovered in West Sussex. On 20 July 1832, fossil collector Gideon Mantell wrote to Professor Benjamin Silliman that when a gunpowder explosion had demolished a quarry rock face in Tilgate Forest, several of the boulders freed showed the bones of a saurian. A local fossil dealer had assembled the about fifty pieces, described by him as a "great consarn of bites and boanes". Having doubts about the value of the fragments, Mantell had nevertheless purchased the pieces and soon discovered they could be united into a single skeleton, partially articulated. Mantell was delighted with the find because previous specimens of "Megalosaurus" and "Iguanodon" had consisted of single bone elements. The discovery in fact represented the most complete non-avian dinosaur skeleton known at the time. He was strongly inclined to describe the find as belonging to the latter genus, but during a visit by William Clift, the curator of the Royal College of Surgeons of England museum, and his assistant John Edward Gray, he began to doubt the identification. Clift was the first to point out that several plates and spikes were probably part of a body armour, attached to the back or sides of the rump. In November 1832 Mantell decided to create a new generic name: "Hylaeosaurus". It is derived from the Greek ὑλαῖος, "hylaios", "of the wood". Mantell originally claimed the name "Hylaeosaurus" meant "forest lizard", after the Tilgate Forest in which it was discovered. Later, he claimed that it meant "Wealden lizard" ("wealden" being another word for "forest"), in reference to the Wealden Group, the name for the early Cretaceous geological formation in which the dinosaur was first found.
Pararhabdodon (meaning "near fluted tooth" in reference to "Rhabdodon") is a genus of tsintaosaurin hadrosaurid dinosaur, from the Maastrichtian-age Upper Cretaceous Tremp Formation of Spain. The first remains were discovered from the Sant Romà d’Abella fossil locality and assigned to the genus "Rhabdodon", and later named as the distinct species Pararhabdodon isonensis in 1993. Known material includes assorted postcranial remains, mostly vertebrae, as well as from the skull. Specimens from other sites, including remains from France, a maxilla previously considered the distinct taxon Koutalisaurus kohlerorum, an additional maxilla from another locality, the material assigned to the genera "Blasisaurus" and "Arenysaurus", and the extensive Basturs Poble bonebed have been considered at different times to belong to the species, but all of these assignments have more recently been questioned.
"Pararhabdodon" has been classified in a number of different positions within Iguanodontia since its remains were first discovered in the mid-1980s. When the first speciemens were originally discovered and described by Casanovas-Cladellas and colleagues, they were not thought to belong a new taxon at all, but merely to belong to the genus "Rhabdodon", and of an uncertain species. Not long after, they would be recognized of those of a unique species, and in 1993 they were given their modern name. At this point it was still thought to be a rhabdodontid or some other type of primitive iguanodont closely related to them. This was based on the characteristics of its vertebrae, as the cranial remains had not yet been found. In 1994 these cranial remains would be found - two maxilla - and these would be used by the original authors to establish it as a species of hadrosaur.
During the 1830s, Danish naturalist Peter Wilhelm Lund and his assistants collected fossils in the calcareous caves near the small town of Lagoa Santa, Minas Gerais, Brazil. Among the thousands of fossils found, he recognized a few isolated cheek teeth as belonging to a hyena, which he named "Hyaena neogaea" in 1839. After more material was found (including canine teeth and foot bones), Lund concluded the fossils instead belonged to a distinct genus of felid, though transitional to the hyenas. He stated it would have matched the largest modern predators in size, and was more robust than any modern cat. Lund originally wanted to name the new genus "Hyaenodon", but realizing this had recently become preoccupied by another prehistoric predator, he instead named it "Smilodon populator" in 1842. He explained the Ancient Greek meaning of "Smilodon" as ("smilē"), a scalpel or two-edged knife, and "οδόντος" ("odontús"), tooth. This has also been translated as "tooth shaped like double-edged knife". He explained the species name "populator" as "the destroyer", which has also been translated as "he who brings devastation". By 1846, Lund had acquired nearly every part of the skeleton (from different individuals), and more specimens were found in neighboring countries by other collectors in the following years. Though some later authors used Lund's original species name "neogaea" instead of "populator", it is now considered an invalid "nomen nudum" ("naked name"), as it was not accompanied with a proper description and no type specimens were designated. Some South American specimens have been referred to other genera, subgenera, species, and subspecies, such as "Smilodontidion riggii", "Smilodon" ("Prosmilodon") "ensenadensis", and "S. bonaeriensis", but these are now thought to be junior synonyms of "S. populator".
Ichthyosaur fossils of the species "Shonisaurus popularis" were first discovered in the area in 1928. Excavations were conducted through the 1960s, and the remains of approximately 40 ichthyosaurs were found. Until 2004, these remains included the largest ichthyosaurs ever discovered. Several specimens were left where they were found, and can be viewed by the public. These specimens are protected from the elements by a large barn. The fossils are about a 10-minute drive from the Berlin ghost town. The ichthyosaur fossils were designated a National Natural Landmark in 1973.
Specimens assigned to "Pteranodon" have been found in both the Smoky Hill Chalk deposits of the Niobrara Formation, and the slightly younger Sharon Springs deposits of the Pierre Shale Formation. When "Pteranodon" was alive, this area was covered by a large inland sea, known as the Western Interior Seaway. Famous for fossils collected since 1870, these formations extend from as far south as Kansas in the United States to Manitoba in Canada. However, "Pteranodon" specimens (or any pterosaur specimens) have only been found in the southern half of the formation, in Kansas, Wyoming, and South Dakota. Despite the fact that numerous fossils have been found in the contemporary parts of the formation in Canada, no pterosaur specimens have ever been found there. This strongly suggests that the natural geographic range of "Pteranodon" covered only the southern part of the Niobrara, and that its habitat did not extend farther north than South Dakota.
"Pteranodon" fossils are known primarily from the Niobrara Formation of the central United States. Broadly defined, "Pteranodon" existed for more than four million years, during the late Coniacian to early Campanian stages of the Cretaceous period. The genus is present in most layers of the Niobrara Formation except for the upper two; in 2003, Kenneth Carpenter surveyed the distribution and dating of fossils in this formation, demonstrating that "Pteranodon sternbergi" existed there from 88 to 85 million years ago, while "P. longiceps" existed between 86 and 84.5 million years ago. A possible third species, which Kellner named "Geosternbergia maysei" in 2010, is known from the Sharon Springs member of the Pierre Shale Formation in Kansas, Wyoming, and South Dakota, dating to between 81.5 and 80.5 million years ago.
Pteranodon (; from Greek πτερόν ( pteron, "wing") and ἀνόδων (anodon, "toothless") is a genus of pterosaur that included some of the largest known flying reptiles, with wingspans over 8 meters (26 feet). They lived during the late Cretaceous geological period of North America in present-day Kansas, Alabama, Nebraska, Wyoming, and South Dakota. More fossil specimens of "Pteranodon" have been found than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved with nearly complete skulls and articulated skeletons. It was an important part of the animal community in the Western Interior Seaway.
The next year, O. C. Marsh led a paleontological expedition into the western United States on behalf of Yale University. Late that November they visited the area around Fort Wallace. Among the fossils found by Marsh's crew in western Kansas were the far ends of two pterosaur wing metacarpals. These were the first scientifically documented fossils of the pterosaur that would later be named Pteranodon. This formal naming occurred six years later, in 1876.