Instruct: Given a question, retrieve passages that can help answer the question.
Query: Where were the first mososaur fossils discovered?
Mosasaurs (from Latin "Mosa" meaning the 'Meuse river', and Greek σαύρος "sauros" meaning 'lizard') are an extinct group of large marine reptiles containing 38 genera in total. Their first fossil remains were discovered in a limestone quarry at Maastricht on the Meuse in 1764. Mosasaurs probably evolved from an extinct group of aquatic lizards known as aigialosaurs in the Early Cretaceous. During the last 20 million years of the Cretaceous period (Turonian-Maastrichtian ages), with the extinction of the ichthyosaurs and pliosaurs, mosasaurs became the dominant marine predators. They became extinct as a result of the K-Pg event at the end of the Cretaceous period, about 66 million years ago.
The Morrison Formation is a sequence of shallow marine and alluvial sediments which, according to radiometric dating, ranges between 156.3 million years old (Ma) at its base, to 146.8 million years old at the top, which places it in the late Oxfordian, Kimmeridgian, and early Tithonian stages of the Late Jurassic period. This formation is interpreted as a semiarid environment with distinct wet and dry seasons. The Morrison Basin where dinosaurs lived, stretched from New Mexico to Alberta and Saskatchewan, and was formed when the precursors to the Front Range of the Rocky Mountains started pushing up to the west. The deposits from their east-facing drainage basins were carried by streams and rivers and deposited in swampy lowlands, lakes, river channels and floodplains. This formation is similar in age to the Solnhofen Limestone Formation in Germany and the Tendaguru Formation in Tanzania. In 1877 this formation became the center of the Bone Wars, a fossil-collecting rivalry between early paleontologists Othniel Charles Marsh and Edward Drinker Cope.
"Pteranodon" was the first pterosaur found outside of Europe. Its fossils first were found by Othniel Charles Marsh in 1870, in the Late Cretaceous Smoky Hill Chalk deposits of western Kansas. These chalk beds were deposited at the bottom of what was once the Western Interior Seaway, a large shallow sea over what now is the midsection of the North American continent. These first specimens, YPM 1160 and YPM 1161, consisted of partial wing bones, as well as a tooth from the prehistoric fish "Xiphactinus", which Marsh mistakenly believed to belong to this new pterosaur (all known pterosaurs up to that point had teeth). In 1871, Marsh named the find ""Pterodactylus oweni"", assigning it to the well-known (but much smaller) European genus "Pterodactylus". Marsh also collected more wing bones of the large pterosaur in 1871. Realizing that the name he had chosen had already been used for Harry Seeley's European pterosaur species "Pterodactylus oweni" in 1864, Marsh re-named his giant North American pterosaur Pterodactylus occidentalis, meaning "Western wing finger," in his 1872 description of the new specimen. He also named two additional species, based on size differences: Pterodactylus ingens (the largest specimen so far), and Pterodactylus velox (the smallest).
Marc Boada discovered, in the late 1990s, a new fossil-bearing locality from the village of Basturs. As the village already lent its name to a locality where dinosaur eggs had been found, it was named the Basturs Poble (BP) locality, "poble" derived from the word for village. The locality belongs to the Conques Formation of the Tremp Group, dated to the late-Early Maastrictian. Over ten years, from 2001 through 2011, field work was done at this site, and a massive bonebed - consisting of some one thousand skeletal elements - was discovered. The discovery was first reported in the literature in an abstract at the "55th Symposium for Vertebrate Palaeontology and Comparative Anatomy and the 16th meeting of the Symposium for Palaeontological Preparation and Conservation" conference in 2007. The authors, then, approached the bonebed under the hypothesis of a single species being present at the site. It was reported that all ontogenetic stages were present. Later, a poster presentation and abstract at another conference suggested the specimens may have belonged to "Koutalisaurus kohlororum", but they cautioned an adult dentary from the bonebed would need to be discovered to test the hypothesis. This was later rejected, as "Koutalisaurus" was found to be an invalid "nomen dubium". In 2018, the material was instead noted as possibly referrable to "Pararhabdodon isonensis".
Three specimens have been assigned to "Sinosauropteryx prima": the holotype GMV 2123 (and its counter slab [opposite face], NIGP 127586), NIGP 127587, and D 2141. Another specimen, IVPP V14202, was assigned to the genus but not to the only species by Zhang and colleagues. The assignment of an additional larger specimen to "S. prima", GMV 2124, was later found to be in error. All of the fossils were found in the Jianshangou or Dawangzhangzi Beds of the Yixian Formation in the Beipiao and Lingyuan regions of Liaoning, China. These fossil beds have been dated to 124.6–122 million years ago, during the early Aptian stage of the Early Cretaceous.
Platecarpus ("flat wrist") is an extinct genus of aquatic lizards belonging to the mosasaur family, living around 84–81 million years ago during the middle Santonian to early Campanian, of the Late Cretaceous period. Fossils have been found in the United States and a possible specimen in Belgium and Africa. A well-preserved specimen of "Platecarpus" shows that it fed on moderate-sized fish, and it has been hypothesized to have fed on squid, and ammonites as well. Like other mosasaurs, it was initially thought to have swum in an eel-like fashion, although another study suggests that it swam more like modern sharks. An exceptionally well-preserved specimen of "P. tympaniticus" known as LACM 128319 shows skin impressions, pigments around the nostrils, bronchial tubes, and the presence of a high-profile tail fluke, showing that it and other mosasaurs did not necessarily have an eel-like swimming method, but were more powerful, fast swimmers. It is held in the Natural History Museum of Los Angeles County.
Alamosaurus (; meaning "Ojo Alamo lizard") is a genus of titanosaurian sauropod dinosaurs, containing a single known species, Alamosaurus sanjuanensis, from the late Cretaceous Period of what is now southern North America. Isolated vertebrae and limb bones indicate that it reached sizes comparable to "Argentinosaurus" and "Puertasaurus", which would make it the largest dinosaur known from North America. Its fossils have been recovered from a variety of rock formations spanning the Maastrichtian age of the late Cretaceous period. Specimens of a juvenile "Alamosaurus sanjuanensis" have been recovered from only a few meters below the Cretaceous-Paleogene boundary in Texas, making it among the last surviving non-avian dinosaur species.
220,000,000 BC, Late Triassic (Arizona) − This episode follows the evolution of the first dinosaurs and other prehistoric creatures in the late Triassic period.152,000,000 BC, Late Jurassic (Colorado) − This episode follows the story of a young female "Diplodocus" and her brothers and sisters in the late Jurassic period.149,000,000 BC, Late Jurassic (Oxfordshire) − This episode follows the life of several baby "Ophthalmosaurus" and a bull "Liopleurodon" in the late Jurassic period.127,000,000 BC, Early Cretaceous (Brazil) − This episode follows a male "Ornithocheirus" on his long journey to the mating grounds in the early Cretaceous period.106,000,000 BC, Early Cretaceous (Antarctica) − This episode follows the hardships of an Antarctican "Leaellynasaura" clan in the early Cretaceous period.65,500,000 BC, Late Cretaceous (Montana) − This episode follows the final years of the dinosaurs and a mother "Tyrannosaurus rex" and her babies in the late Cretaceous period.49,000,000 BC, Early Eocene (Germany) − This episode follows the life of a mother "Leptictidium" and her babies in the dense forests of Germany in the early Eocene epoch.36,000,000 BC, Late Eocene (Pakistan) − This episode follows the sea journey of a pregnant female "Basilosaurus" search for food in the late Eocene epoch.25,000,000 BC, Late Oligocene (Mongolia) − This episode follows the life of a young "Paraceratherium" and his mother from his birth in the late Oligocene epoch.3,200,000 BC, Late Pliocene (Ethiopia) − This episode follows the hardships of an "Australopithecus" group and a young called Blue in the late Pliocene epoch.1,000,000 BC, Early Pleistocene (Paraguay) − This episode follows a male "Smilodon" called Half-Tooth after he lost his territory in the early Pleistocene epoch.30,000 BC, Late Pleistocene (North Sea, Belgium, and Swiss Alps) − This episode follows a herd of woolly mammoth journey to the Alps in the last Ice Age of the late Pleistocene epoch.This episode details a list of animals from the first season that the developers thought were the most impressive creatures on Earth.
Recent molecular phylogenetic studies suggest that most placental orders diverged late in the Cretaceous period, about 100 to 85 million years ago, but that modern families first appeared later, in the late Eocene and early Miocene epochs of the Cenozoic period. Fossil-based analyses, on the contrary, limit the placentals to the Cenozoic. Many Cretaceous fossil sites contain well-preserved lizards, salamanders, birds, and mammals, but not the modern forms of mammals. It is likely that they simply did not exist, and that the molecular clock runs fast during major evolutionary radiations. On the other hand, there is fossil evidence from of hoofed mammals that may be ancestors of modern ungulates.
Fossils of "Smilodon" were discovered in North America from the second half of the 19th century onwards. In 1869, American paleontologist Joseph Leidy described a maxilla fragment with a molar, which had been discovered in a petroleum bed in Hardin County, Texas. He referred the specimen to the genus "Felis" (which was then used for most cats, extant as well as extinct) but found it distinct enough to be part of its own subgenus, as "F." ("Trucifelis") "fatalis". The species name means "fate" or "destiny", but it is thought Leidy intended it to mean "fatal". In an 1880 article about extinct American cats, American paleontologist Edward Drinker Cope pointed out that the "F. fatalis" molar was identical to that of "Smilodon", and he proposed the new combination "S. fatalis". Most North American finds were scanty until excavations began in the La Brea Tar Pits in Los Angeles, where hundreds of individuals of "S. fatalis" have been found since 1875. "S. fatalis" has junior synonyms such as "S. mercerii", "S. floridanus", and "S. californicus". American paleontologist Annalisa Berta considered the holotype of "S. fatalis" too incomplete to be an adequate type specimen, and the species has at times been proposed to be a junior synonym of "S. populator". Swedish paleontologists Björn Kurtén and Lars Werdelin supported the distinctness of the two species in 1990.
While the first "Pteranodon" wing bones were collected by Marsh and Cope in the early 1870s, the first "Pteranodon" skull was found on May 2, 1876, along the Smoky Hill River in Wallace County (now Logan County), Kansas, USA, by Samuel Wendell Williston, a fossil collector working for Marsh. A second, smaller skull soon was discovered as well. These skulls showed that the North American pterosaurs were different from any European species, in that they lacked teeth and had bony crests on their skulls. Marsh recognized this major difference, describing the specimens as "distinguished from all previously known genera of the order Pterosauria by the entire absence of teeth." Marsh recognized that this characteristic warranted a new genus, and he coined the name "Pteranodon" ("wing without tooth") in 1876. Marsh reclassified all the previously named North American species from "Pterodactylus" to "Pteranodon". He considered the smaller skull to belong to "Pteranodon occidentalis", based on its size. Marsh classified the larger skull, YPM 1117, in the new species "Pteranodon longiceps", which he thought to be a medium-sized species in between the small "P. occidentalis" and the large "P. ingens". Marsh also named several additional species: Pteranodon comptus and "Pteranodon nanus" were named for fragmentary skeletons of small individuals, while "Pteranodon gracilis" was based on a wing bone that he mistook for a pelvic bone. He soon realized his mistake, and re-classified that specimen again into a separate genus, which he named "Nyctosaurus". "P, nanus" was also later recognized as a "Nyctosaurus" specimen.
The Early Cretaceous Epoch spans from 145 million to 100 million years ago. The Early Cretaceous saw the expansion of seaways, and as a result, the decline and extinction of sauropods (except in South America). Many coastal shallows were created, and that caused ichthyosaurs to die out. Mosasaurs evolved to replace them as apex species of the seas. Some island-hopping dinosaurs, like "Eustreptospondylus", evolved to cope with the coastal shallows and small islands of ancient Europe. Other dinosaurs, such as "Carcharodontosaurus" and "Spinosaurus", rose to fill the empty space that the Jurassic-Cretaceous extinction had created. Of the most successful would be the "Iguanodon" which spread to every continent. Seasons came back into effect and the poles grew seasonally colder. Dinosaurs such as the "Leaellynasaura" inhabited the polar forests year-round, while many dinosaurs, such as the "Muttaburrasaurus", migrated there during summer . Since it was too cold for crocodiles, it was the last stronghold for large amphibians, such as the "Koolasuchus". In this epoch Pterosaurs reached their maximum diversity and grew larger, as species like "Tapejara" and "Ornithocheirus" took to the skies. The first true birds evolved, possibly sparking competition between them and the pterosaurs.
The museum's five rooms shows paleontological finds from the Besano area, including the "Besanosaurus", a type of ichthyosaur from the middle Triassic period.
The 6 meter "Besanosaurus leptorhynchus" discovered in 1993 is the biggest sea reptile fossil found in Italy, and was probably a pregnant female.
Since 2001 the museum has also housed the remains of a "Saltriosaurus", discovered in a quarry in Saltrio, which is the first big carnivorous dinosaur found in Lombardia.
Dimetrodon ( or , meaning "two measures of teeth") is an extinct genus of synapsids that lived during the Cisuralian (Early Permian), around 295–272 million years ago (Ma). It is a member of the family Sphenacodontidae. The most prominent feature of "Dimetrodon" is the large neural spine sail on its back formed by elongated spines extending from the vertebrae. It walked on four legs and had a tall, curved skull with large teeth of different sizes set along the jaws. Most fossils have been found in the southwestern United States, the majority coming from a geological deposit called the Red Beds of Texas and Oklahoma. More recently, fossils have been found in Germany. Over a dozen species have been named since the genus was first described in 1878.
In 1983, a relatively complete skeleton was excavated from the Smokejacks pit in Surrey, Engand. These remains were described by British paleontologists Alan J. Charig and Angela C. Milner in 1986 as the holotype of a new species, "Baryonyx walkeri." After the discovery of "Baryonyx," many new genera have since been described, the majority on very incomplete remains. However, other finds bear enough fossil material and distinct anatomical features to be assigned with confidence. Paul Sereno and colleagues described "Suchomimus" in 1998"," a baryonychine from Niger on the basis of a partial skeleton found in 1997. Later in 2004, partial jaw bones were recovered from the Alcântara Formation, these were referred to a new genus of spinosaurine named "Oxalaia" in 2011 by Alexander Kellner.
Among the first scientists to explore the Muirkirk Deposit was Maryland state geologist Phillip Thomas Tyson. He brought some of the strange bones discovered in the iron mines to a meeting of the Maryland Academy of Sciences in 1859, where his colleagues identified them as dinosaurs. Paleontologist Othniel Charles Marsh was also interested in Maryland fossils. In the winter of 1887, he sent John Bell Hatcher to search the iron mines. Hatcher recovered hundreds of fossils, including the remains of ancient turtles and crocodiles. In the 1890s, Smithsonian Institution scientists Charles Gilmore and Arthur Bibbins also visited Prince George's County, uncovering dinosaur teeth and other fossils that were added to the Smithsonian collection.
Paleontology has a long history in the State of Illinois, stretching at least as far back as the 1850s, when the first Mazon Creek fossils were being found. Early in the ensuing 20th century a Silurian age fossil reef system was discovered in the state. In the 1950s Francis Tully discovered the "monster" that would later be named in his honor. The Pennsylvanian species "Tullimonstrum gregarium" ("Tully Monster") is the Illinois state fossil. It is one of the few officially designated state fossils that is endemic to the state it represents.
Many important fossils have been found in the sandstones and clays of the Weald, including, for example, "Baryonyx". The famous scientific hoax of Piltdown Man was claimed to have come from a gravel pit at Piltdown near Uckfield. The first "Iguanodon" was identified after a Mary Mantell unearthed some fossilised teeth by a road in Sussex in 1822. Her husband, Gideon Mantell, noticed they were similar to modern iguana teeth but many times larger; this important find led to the discovery of dinosaurs.
"Sinosauropteryx" was a small theropod with an unusually long tail and short arms. The longest known specimen reaches up to in length, with an estimated weight of It was a close relative of the similar but older genus "Compsognathus", both genera belonging to the family Compsognathidae. Only one species of "Sinosauropteryx" has been named: "S. prima", meaning "first" in reference to its status as the first feathered non-avialian dinosaur species discovered. Three specimens have been described. The third specimen previously assigned to this genus represents either a second, as-yet unnamed species or a distinct, related genus.
Controversy regarding the identity of the filaments preserved in the first "Sinosauropteryx" specimen began almost immediately, as the team of scientists spent three days in Beijing examining the specimen under a microscope. The results of their studies (reported during a press conference at the Philadelphia Academy of Natural Sciences on Thursday, April 24, 1997) were inconclusive; the team agreed that the structures preserved on "Sinosauropteryx" were not modern feathers, but suggested further research was required to discover their exact nature. Palaeontologist Alan Feduccia, who had not yet examined the specimen, wrote in "Audubon Magazine" that the structures of "Sinosauropteryx" (which he considered at the time to be a synonym of "Compsognathus", as "Compsognathus prima") were stiffening structures from a frill running along the back, and that dinosaur palaeontologists were engaging in wishful thinking when equating the structures with feathers. Subsequent publications saw some of the team members disagreeing over the identity of the structures.
Prieto-Márquez returned again to the dentary in 2013, in a study alongside colleagues providing a review and investigation of hadrosaurs from all over Europe. Further preparation of the specimen in the time since his last study regarding it revealed the uniqueness of the dentary had been exaggerated significantly by reconstruction of the specimen when it was first prepared in the 1990s. The "Tsintaosaurus" specimens showing the similar condition were found to have been distorted from a similar process. Correcting for the innaccuracies, the Los Llaus dentary is indistinguishable from that of multiple lambeosaurines, and shares no particular connection to "Tsintaosaurus". With this, their reasoning for assignment of the specimen to "Pararhabdodon" was voided, and the specimen is now considered a completely indeterminate lambeosaurine dentary.
One of the first major fossil finds of the 20th century in Pennsylvania was the 1902 discovery of dinosaur tracks at a fisher's Quarry near Graterford in Montgomery County. Another prominent early 1900s discovery was a lower jaw from the amphibian "Calamops" was discovered in the Late Triassic Stockton Formation at Holicong. This is the oldest vertebrate fossil from either the Hartford or Newark Basins. In 1923 the Reading Railroad tunnel at Gwynned was converted into an open cut, exposing many Late Triassic fossils. Later, in 1933, two "Anchisauripus" tracks were discovered near Yocumtown in York County. Dinosaur tracks dating back to the Late Triassic were discovered near the Gettysburg battle sites in 1933.
It was named after Morrison, Colorado, where the first fossils in the formation were discovered by Arthur Lakes in 1877. That same year, it became the center of the Bone Wars, a fossil-collecting rivalry between early paleontologists Othniel Charles Marsh and Edward Drinker Cope. In Colorado, New Mexico, and Utah, the Morrison Formation was a major source of uranium ore.
Additional remains of "Simosaurus" were found in Franconia and Württemberg in Germany. Duke William of Württemberg discovered a complete skull and sent it to von Meyer in 1842. Von Meyer named "S. guilelmi" on the basis of this skull, noting that it was smaller and narrower than those of the type species. A complete skeleton first referred to "S. guilelmi" has been designated the neotype of "Simosaurus". Some German fossils have been found in the stratigrafically younger Keuper deposits, but are very rare. "Simosaurus" is present in biozones of the Muschelkalk that are distinguished by different ammonite fauna. "Simosaurus" first appears in the "nodosus" biozone, where fossils of the ammonite "Ceratites nodosus" are abundant. Specimens becomescommon in the slightly younger "dorsoplanus" biozone, characterized by the ammonite "Ceratites dorsoplanus".
Infant and juvenile fossils of "Mussaurus" were first discovered by an expedition led by Jose Bonaparte during the 1970s to the El Tranquilo Formation. There the team found fossilized eggs and hatchlings, which added insight into the reproductive strategies of "Mussaurus" and other sauropodomorph dinosaurs. The first adult specimens of "Mussaurus" were described in 2013. Some of these specimens had first been described in 1980 and were originally attributed to the genus "Plateosaurus".