Ethologists usually study how animals behave in their natural environment.
Ethologists study animal behavior.
Animal behaviorists usually study how animals behave in their natural environment.
What usually studies how animals behave in their natural environment?

a: psychologists
b: zookeepers
c: looseness of dirt or compost
d: it's state
e: that material's properties
f: teachers
g: sensory neurons
h: animal behaviorists