The genetic richness of "Testudo graeca" is also shown in its crossbreeding. "Testudo marginata" is also closely related to the Greek or common tortoise ("Testudo graeca"). In the spur-thighed tortoise ("Testudo graeca"), both males and females have spurs. The Tunisian tortoise or Nabeul tortoise ("Testudo graeca nabeulensis") is a subspecies of Greek tortoises. Like the golden eagle, the eagle-owl may attack spur-thighed tortoises ("Testudo graeca"). Reptiles native to the region include spur-thighed tortoise ("Testudo graeca buxtoni") and Horsfield's tortoise. The Greek tortoise ("Testudo graeca ibera") is often confused with Hermann's tortoise ("Testudo hermanni" ).