The Munda languages are also polysynthetic and agglutinating. Karaim morphology is suffixing and highly agglutinating. Aymara is an agglutinating and, to a certain extent, a polysynthetic language. Santali, like all Munda languages, is a suffixing agglutinating language. The supernatant fluid is tested for agglutinating activity for guinea pig erythrocytes. Typologically, Estonian represents a transitional form from an agglutinating language to a fusional language. Udi is agglutinating with a tendency towards being fusional. Verb morphology can be described as either fusional or agglutinating depending on the analysis. Ho-Chunk is an agglutinating and somewhat fusional language. Inflected and agglutinating languages may have extremely complicated systems of morphophonemics. Zo is monosyllabic, partially agglutinating tone language. Achylectins, isolated from "Tachypleus tridentatus", show specific agglutinating activity against human A-type erythrocytes. The language is of the agglutinating type with comparatively little productive morphology. Middle Mongol is an agglutinating language that makes nearly exclusive use of suffixes. Voice and aspect are also indicated by means of conjugation, and possibly agglutinating auxiliary verbs.