The olfactory system, or sense of smell, is the sensory system used for smelling (olfaction). Damage to the olfactory bulb results in a loss of olfaction (the sense of smell). Lizards make use of their senses of sight, touch, olfaction and hearing like other vertebrates. Olfaction, taste, and trigeminal receptors (also called chemesthesis) together contribute to flavor. Antennae are the primary organs of olfaction (smell) in Lepidoptera. In the laboratory, cattle can be trained to recognise conspecific individuals using olfaction only. The parahippocampal gyrus houses the topographical map for olfaction. Taste perception is a combination of oral somatosensation and retronasal olfaction. Crocodiles close their nostrils when submerged, so olfaction underwater is unlikely. The stimuli are recognized by the region of the human brain which handles olfaction. Lemur communication can be transmitted through sound, sight, and smell (olfaction). Cannabinoids are not known to have effects on olfaction in humans. Humans can detect blood relatives from olfaction. Another major function of the nose is olfaction, the sense of smell. It is the senses of touch and olfaction that are most important.