Typhlosoles occur in bivalve mollusks, lampreys and some annelids and echinoderms. Crustaceans, nematodes, trematodes, and polychaete annelids also serve as parasites. The gregarines are generally parasites of annelids, arthropods, and mollusks. Other invertebrates including insect larvae, annelids and arachnids possess bioluminescent abilities. Compound eyes are common in arthropods, annelids and some bivalved molluscs. In saline regions, there are abundant polychaete annelids, amphipods, and some mollusks such as clams. This hypothesis groups annelids with molluscs and brachiopods in another superphylum, Lophotrochozoa. The Lophotrochozoa includes the molluscs, annelids, brachiopods, nemerteans, bryozoa and entoprocts. Nociceptors have been found in nematodes, annelids and mollusks. Eibye-Jacobsen regarded bristles as a feature shared by molluscs, annelids and brachiopods. Most nemerteans are carnivores, feeding on annelids, clams and crustaceans. Hox gene expression has also been studied in brachiopods, annelids, and a suite of molluscs. The carnivores feed on annelids, crustaceans, and fish. Some annelids and bivalves also have apposition eyes. Evidence of the relationship between molluscs, annelids and lophophorates was found in 1995.