Mitochondria are essential organelles in eukaryotic cells. Corneocytes are keratinocytes without nuclei and cytoplasmic organelles. The vacuoles shrink and other cellular organelles proliferate. Vacuoles are large intracellular storage organelles in plants cells. Eukaryotic cells have nucleus with multiple organelles and more DNA arranged in linear chromosomes. Chloroplasts are one of many types of organelles in the plant cell. The earliest evidence of complex eukaryotes with organelles (such as mitochondria) dates from . All dinoflagellates contain red algal plastids or remnant (nonphotosynthetic) organelles of red algal origin. It goes through destroying and recycling the cytoplasmic organelles and macromolecules. Not all eukaryotic cells have each of the organelles listed below. This excludes any part of the cytoplasm that is contained within organelles. The cytosol is thus a liquid matrix around the organelles. Retrograde transport shuttles molecules/organelles away from axon termini toward the cell body. Some protists have various other microtubule-supported organelles. Mature lens fibers have no organelles or nuclei.