The xylem typically lies adaxial with phloem positioned abaxial. The pollen is deposited on the abaxial (off-axis) surface of the stylus. The secondary venation is parallel and the abaxial leaf surfaces are glabrous. Leaves have pointed ends and are densely silvery or golden sericeous on the abaxial side. Plants have five stamens with the abaxial stamen often spurred at the base. The abaxial faces of the laminae are glabrous or moderately hairy. The corolla lobes show glandular hairs at the abaxial surface. The abaxial leaf surface has a spongy white texture. The sporangia are described as round in abaxial view, and oval in lateral view. The abaxial surface (underside) of the leaf is somewhat hairy. The abaxial surfaces of leaf blades are glabrous or scabrous and occasionally pubescent or puberulent.