Some antioxidants include phenolic acids, flavonoids, proanthocyanidins, and anthocyanin. The anthocyanins are subdivided into the sugar-free anthocyanidin aglycones and the anthocyanin glycosides. The upper surface of these lobes contains red anthocyanin pigments and its edges secrete mucilage. The increase in anthocyanin production is accompanied by a decrease in chlorophyll production. Castavinols are another class of colorless molecules derived from colored anthocyanin pigments. The purple colour common in these carrots comes from anthocyanin pigments. The color of purple varieties is due to various anthocyanin pigments. Ruby is a MYB transcriptional activator of genes that produce anthocyanin in citrus fruits. The M gene determines the production of a purple pigment, a type of anthocyanin. The red pigment is identified as the Perilla anthocyanin, "shisonin". The color of purple skin cultivars is due to the anthocyanin nasunin. When this happens, the sugar-breakdown process changes, leading to the production of anthocyanin pigments. Many plants or plant parts contain chemicals from the naturally colored anthocyanin family of compounds. Yellow raspberries and others with pale-colored fruits are lower in anthocyanin content. It has been shown to increase the rate of ethylene production and anthocyanin concentrations.