In these females, copulations after the first occur after longer courtships. Feeding by the dragonet occurs throughout the day, including the intervals between courtships and spawning. This dewlap is used in courtships and territorial displays. In it, Halkett gives a detailed account of her courtships and marriage. These courtships take place during spring migration and shortly after arriving on the breeding grounds. Long-term strategies are characterized by extended courtships, high investment, and few sexual partners. As a result, they must attempt to elicit courtships to maximize their reproductive options. For low-intensity courtships, males and females walk in unison with their heads raised.