Each one of the four haploid nuclei migrates into its own basidiospore. "Ganoderma" can be differentiated from other polypores because they have a double-walled basidiospore. He also observed basidiospore development in boletoid and tricholomatoid fungi. "Coprinopsis radiata" has been used as a model organism to study basidiospore formation. In these cases the basidiospore typically lacks a hilar appendage, and no forcible discharge occurs.