This includes kangaroos, wallabies, koalas, platypus, echidna, emu, kookaburras and dragon lizards. The other mascots were Millie the Echidna and Syd the Platypus. Platypuses and two genera of echidna are Australian monotremes. Both the platypus and echidna species have spurs on their hind limbs. A patella is also present in the living monotremes, the platypus and the echidna. Numerous physiological adaptations aid the lifestyle of the short-beaked echidna. The five subspecies of the short-beaked echidna are each found in different geographical locations. The musculature of the short-beaked echidna has a number of unusual aspects. The player character is guided and instructed by the voice of Tikal the Echidna. She appears to have a crush on Knuckles the Echidna. "Sonic 3" features the debut of Sonic's rival, Knuckles the Echidna. Next comes the Chimera (whose mother is unclear, either Echidna or the Hydra). This immense serpent, a child of Typhon and Echidna, guarded the Golden Fleece at Colchis. Kathie Lee Gifford was also present as the voice of Typhon's mate, Echidna. He then found the dracaena of Scythia (sometimes identified as Echidna) in a cave.