Manus and pes range from plantigrade to digitigrade. Their method of locomotion ranged from plantigrade to digitigrade. A plantigrade foot is the primitive condition for mammals; digitigrade and unguligrade locomotion evolved later. They are quadrupedal and, when on the ground, they walk digitigrade and plantigrade. Whereas the aardvark is considered digitigrade, it appears at time to be plantigrade. Like this archosaur, basal pterosaur lineages have plantigrade hindlimbs that show adaptations for saltation. Terrestrial animals using their soles for locomotion are called plantigrade. Embrithopods were rhinoceros-like herbivorous mammals with plantigrade feet, and desmostylians were hippopotamus-like amphibious animals. Humans usually walk with the soles of their feet on the ground, in plantigrade locomotion. The paws have naked and padded soles and have a plantigrade gait. Unlike most other land carnivorans, bears are plantigrade. "Dinictis" walked plantigrade (flat-footed), unlike modern felids. The manus and pes are plantigrade or subplantigrade. It has plantigrade, pentadactyl feet with two phalanges. In contrast, most caniforms are terrestrial, have non-retractile claws and tend to be plantigrade.