The pseudo-elaters are multi-cellular, unlike the elaters of liverworts. Glebal elaters are aseptate and not branched. Other names include elaters, snapping beetles, spring beetles or skipjacks. Under moist conditions, the elaters curl tightly around the spore. The elaters are yellowish, often branches, and varying in size and form. In the liverworts, elaters are cells that develop in the sporophyte alongside the spores. Elaters come in a variety of forms, but are always associated with plant spores. In the horsetails, elaters are four ribbon-like appendages attached to the spores.