The anthers are basifixed (attached at their base). The anthers are basifixed, as in all of Hamamelidaceae. The anthers are basifixed, and the leaves are arranged in whorls. Submerged leaves bearing sori that are surrounded by basifixed membranous indusia (sporocarps). The flower pistil is 5 celled and the anthers dehiscing via pores and basifixed.