Sori are produced in groups of one to three on fertile pinnule veins. The sori are circular and occur on either side of the fertile pinnule midvein. Sori are circular and occur in one to three rows along the pinnule midvein. Sori are produced in four to six pairs along the pinnule midvein. Sori are round, occur near the fertile pinnule midvein. Two to four sori occur per fertile pinnule. Fertile pinnule, bearing several tufts of microsporangia, magnified. The second pinnule is massive, hard and curved. Sori are close to fertile pinnule midveins and covered by thin indusia. The terminal pinnule of each frond was oval or oblong in shape.