Anthropology expanded to include paleoanthropology and primatology. He draws evidence from evolutionary biology, moral and political philosophy, primatology, linguistics, and anthropology. Her chief areas of research are in ornithology and primatology. He specialises in the fields of primatology, herpetology, biodiversity and conservation of tropical forests. Primatology is the scientific study of primates. Her works have sparked debate in primatology, philosophy, and developmental biology. The intermembral index is used frequently in primatology, since it helps predict primate locomotor patterns. The study of primatology looks at the biological and psychological aspects of non-human primates. The discipline of Japanese primatology was developed out of animal ecology. Japanese primatology is a carefully disciplined subjective science. Western primatology stems primarily from research by North American and European scientists. Early field primatology tended to focus on individual researchers. There are two main centers of primatology, Western primatology and Japanese primatology. For Japanese researchers in primatology, the findings of the team are emphasised over the individual. The award is given every two years for distinguished contributions to primatology.