Professor Asen I. Hadjiolov was a well-known Bulgarian histologist and cytologist. These glands are named after Victor von Ebner, an Austrian histologist. Jones attained a reputation as a histologist and as a clinical observer. David M. Sever is an American herpetologist, histologist, anatomist and reproductive biologist. John Thomas Quekett (1815 - 1861) was a pioneering histologist. Sir William Bowman, histologist, ophthalmologist and surgeon, was born there in 1816. His brother was the histologist Nevill Willmer, while another brother, Gordon Willmer, was a judge.