Their principal predators are Nile crocodiles, leopards, lions and hyenas. They may be preyed on by lions, leopards, spotted hyenas and African wild dogs. Lions defend their kills from scavengers such as vultures and hyenas. A separate area hosts zebras, impalas, kudu, hyenas, and baboons. Cubs also die from starvation and abandonment, and predation by leopards, hyenas and wild dogs. Predators of the greater kudu generally consist of lions, Spotted hyenas, and African hunting dogs. There are records of some male leopards preying on hyenas. The most frequently observed animals are giraffes, cape buffalo, hyenas, and lions. Caracals, hyenas and servals attack calves. The young are occasionally taken by hyenas and lions. The feliforms include families such as the cats, the hyenas, the mongooses and the civets. Red foxes may compete with striped hyenas on large carcasses. The onager is preyed upon by apex predators such as Persian leopard and striped hyenas. The gnolls are a race of humanoid hyenas. Cases of African wild dogs scavenging from spotted hyenas are rare.