North American felid predators of red foxes include cougars, Canadian lynxes and bobcats. Felis lunensis (Martelli's cat) is an extinct felid of the subfamily Felinae. It is not a formal cat breed, but a landrace of felid hybrids. The cougar ("Puma concolor") is a large felid of the subfamily Felinae. Leptofelis is an extinct genus of "Pseudaelurus"-grade felid found in Spain. Unlike other felid species, margays are not induced ovulators. The jaguar is the largest felid that is indigenous to Nicaragua. Mortality in affected felid litters varies between 20 and 100%. Carnassial, the Felid (a type of proto-cat) also has a hidden desire. The first felid, "Proailurus", originated in Asia during the late Oligocene and spread to Europe. A member of this family is called a felid. Modelling of felid coat pattern transformations revealed that nearly all patterns evolved from small spots. A member of this family is also called a felid. Felid forelimb development during ontogeny (changes during growth) has remained tightly constrained.