However, it fills the purpose as the female counterpart of the male spermatid in spermatogenesis. In the spermatid stage, the sperm develops the familiar tail. Meiosis I. This is observed by a reduction in spermatid presence in "repro4" mutants. Spermatid DNA also undergoes packaging, becoming highly condensed. The spermatid is the haploid male gametid that results from division of secondary spermatocytes. ICAM-2 molecules regulate spermatid adhesion on the apical side of the barrier (towards the lumen).