Later studies have sought further evidence of unique characteristics linking the megabat and primate brains. The proportions are equivalent the a smaller megabat or larger microbat of modern chiropteran fauna. It is the only megabat that is endemic to Fiji. A 2004 review stated that 28% of megabat species weigh less than . Notable terrestrial species are the Pacific boa and the Flying Megabat, which has a wingspread. As of 2019, the IUCN had evaluations for 187 megabat species. In Australia, five genera and eight species of megabat are present. Culling campaigns can dramatically reduce megabat populations. Many megabat species are highly gregarious or social. It forms harmonious mixed species colonies with another megabat, the large flying fox ("Pteropus vampyrus"). The Egyptian fruit bat is considered a medium-sized megabat. As of 2018, 197 species of megabat had been described. A 1992 summary of forty-one megabat genera noted that twenty-nine are tree-roosting genera. Wahlberg's epauletted fruit bat ("Epomophorus wahlbergi") is a species of megabat in the family Pteropodidae. Megabat social behavior includes using sexual behaviors for more than just reproduction.