Other organisms eaten include other fly larvae, insect eggs, rotifers and testate amoebae. "Difflugia" are testate amoebae with an agglutinate shell. Lecythium is a genus of testate amoebae within the Chlamydophryidae. Foraminifera are closely related to testate amoebae. Fossils of arcellinid testate amoebae date back to the Cryogenian period. Dame Ursula died testate in 1555. Euglyphid testate amoebae are closely related to the Foraminifera. The genus Difflugia is the oldest and most diverse of the testate amoebae. Where the deceased dies leaving a valid will, the rules of testate succession apply. The genus "Difflugia" is the oldest and most diverse of the testate amoebae. Legatees are beneficiaries in testate succession only. Two cladoceran, two ostracod and one testate amoeba species were originally discovered in Licancabur Lake. Fossils of testate amoeba (or Arcellinida) first appear during the Cryogenian period. Species of Amoebozoa may be either shelled (testate), or naked, and cells may possess flagella. The Will was published testate later that year on December 19 1916.