There was considerable debate whether pterosaurs ambulated as quadrupeds or as bipeds. The dinosaurs are classified in 6 families&colon; large carnivorous, armored quadrupeds, light bipeds, etc. Other prosimians are arboreal quadrupeds and climbers. Monboddo cast early humans as wild, solitary, herbivorous quadrupeds. Most monkeys are both arboreal and terrestrial quadrupeds and climbers. Not all tetrapods are quadrupeds and not all quadrupeds are tetrapods. Bursas are carnivorous quadrupeds native to Naboo. Although the forelimbs are much smaller than the hind limbs, all aetosaurs were quadrupeds. Howlers are carnivorous, reptilian quadrupeds that may be native to Yavin 4. However, sloths are generally regarded as quadrupeds. Folio 5 verso depicts quadrupeds, livestock, wild beasts, and the concept of the herd. Aphanosauria is a Triassic group of gracile carnivorous quadrupeds which was recognized in 2017. Gesner placed all the species within four general categories&colon; quadrupeds, birds, fish and serpents. As massive quadrupeds, sauropods developed specialized "graviportal" (weight-bearing) limbs. Gualamas are docile and swift quadrupeds native to Naboo.