Its distribution is a factor of climatic, edaphic, or other environmental attributes. They present various topographical and edaphic regions. Organic carbon is vital to soil capacity to provide edaphic ecosystem services. The sixteen acacia species vary over this range, their distribution determined by edaphic conditions. Some small edaphic species are pale yellowish. Due to diverse topographical and edaphic factors, the reserve harbors varied vegetation types. The geographical isolation and special edaphic conditions helped to preserve it too. Edaphic means pertaining to the soil. Market ideotype, climatic ideotype, edaphic ideotype, stress ideotype and disease/pest ideotypes are its other concepts. Among the natural have been listed climatic, edaphic (soil), topographic, biotic and pyric (fire). "Trigonopterus" species inhabit primary tropical forests, both on foliage and edaphic in the litter layer. This largely reflects edaphic drought associated with rapid drainage on the steep limestone topography. Small marble-sized, iron-bearing granules are present at all sites and are likely an edaphic requirement.