The simplest forms of haustoria are small spheres. Some parasitic fungi form haustoria that function in absorption within the host cells. They infect and feed on the plant via haustoria. Urediospores germinate and form haustoria that penetrates into the leaves. The haustoria are less than one millimetre long. Through the establishment of haustoria the pathogen develops an intimate relationship with the host. Inside the host, the fungi will produce hyphae and another specialized structure called haustoria. Larger (usually botanical, not fungal) haustoria do this at the tissue level. Pruning infested branches is not generally effective because the haustoria can infiltrate deeply. The evolutionary event which gave rise to parasitism in plants was the development of haustoria. Fungi in all major divisions form haustoria. "Blumeria" differs from "Erysiphe" in its digitate haustoria and in details of the conidial wall. It draws nutrients from the plant via haustoria that penetrate the epidermal layer. Haustoria arise from intercellular hyphae, appressoria, or external hyphae. As the penetration peg grows between cells, haustoria grow into the cells to gain nutrients.