Ablestar was a liquid rocket stage burning hypergolic propellants fed from gas-pressurized propellant tanks. The Dnepr was a three-stage rocket using storable hypergolic liquid propellants. These engines used toxic hypergolic propellants that required special handling precautions. This was reportedly changed to a hypergolic nitrogen-tetroxide/hydrazine propulsion system. It is a hypergolic propellant in combination with a hydrazine-based rocket fuel. TEMED can also be a component of hypergolic propellants. The Volvo Flygmotor group also used a hypergolic propellant combination. These were the only part of the Saturn V that used hypergolic propellants. Although larger hypergolic engines in some launch vehicles use turbopumps, most hypergolic engines are pressure-fed. Soviet rocket engine researcher Valentin Glushko experimented with hypergolic fuel as early as 1931. The R-4D requires no igniter as it uses hypergolic fuel. Primary among these was that the Ajax fuels were "hypergolic", igniting on contact. This was an efficient hypergolic rocket that was throtteable. The hypergolic combination has an ISP of 322 seconds. Hypergolic rockets are still widely used in upper stages when multiple burn-coast periods are required.