They also tend to live in arid steppe climates. "P. spelaea" inhabited open environment such as mammoth steppe and boreal forest. They inhabit open habitats, especially steppe and savannah. Nomadic pastoralism developed in the Pontic-Caspian steppe beginning in the Chalcolithic. Neolithic peoples of the Eastern European steppe. The majority of people in the steppe are nomads called Mongolians. Corsac foxes seem to only outcompete red foxes in semi-desert and steppe areas. Woolly rhinoceroses mostly fed on grasses and sedges that grew in the mammoth steppe. In addition to these problems, the struggle with the nomads from the northern steppe continued. Mapuches came to practise horse husbandry in the northern part of the Patagonian steppe. The horse was domesticated in the Pontic steppe around 4000 BC. Its ecosystem is complex with unique biodiversity with flora of the Central Asian steppe. A few burrowing species are native to open steppe land. It is associated with the formation of the mammoth steppe and continental glaciation. The modern European bison is likely to have arisen from the steppe bison.