Leaves consist of a long petiole and a rachis. All species have spines on the rachis and petiole. The petiole connects the lead base with the rachis. The pinnae are distinctly alternate along the rachis. Glands are usually present on the rachis and the upper side of the petiole. Their shorter rachis segments lead to spikelets packed closer together. Each grape berry contains a pedicel which attaches to the rachis. Spines may be present on the midrib of leaflets and the rachis of compound leaves. It has a broad midrib (rachis). The flowers are borne in panicles, clustered along the rachis. The leaflets are sessile, directly attached to the rachis without a petiolule. The rachis and attached veins make up the spathe. A pollen cone consists of a slender rachis with numerous spirally arranged microsporophylls around it. Bristles are stiff, tapering feathers with a large rachis but few barbs. This resulting feather is one with a tuft of branched barbs without a rachis.