<P> Luddington has worked mainly in the United States . In February 2011 , the Daily Mail described her as `` on the path to fame and fortune after being cast as Catherine Middleton in William & Kate : The Movie , a TV movie about the relationship between Prince William and Catherine `` Kate '' Middleton . Luddington joined the cast of the Showtime comedy - drama series Californication for its fifth season , playing a nanny . She also joined the cast of season five of the HBO vampire drama True Blood , as Claudette Crane , a faerie . In July 2012 , Luddington joined the cast of Grey 's Anatomy as Dr. Jo Wilson in a recurring role . In June 2013 it was announced that she would be a series regular from season ten onward . In October 2012 , Camilla appeared on a Halloween special of E ! 's popular fashion programme , Fashion Police , alongside a panel of Joan Rivers , Kelly Osbourne , George Kotsiopoulos and Kris Jenner . </P>
when does jo come in grey 's anatomy
July 2012