nature traced herodotus died ''c'' bc described one earliest accounts mutualism observation natural dentistry basking nile crocodiles noted would open mouths give sandpipers safe access pluck leeches giving nutrition sandpiper oral hygiene crocodile ref name egerton aristotle early influence philosophical development ecology student theophrastus made extensive observations plant animal migrations biogeography physiology behavior giving early analogue modern concept ecological niche ref name hughes ref name hughes div
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