With a seasonal harvest of between 1,000 and 1,500 animals, in 1974 the estimated size of the population distributed in the Caucasus region was around 20,000 animals and the density was four animals per square kilometer (10 animals per square mile).
Cunha colleita estacional de entre 1 000 e 1 500 animais, en 1974 o tamaño estimado da poboación distribuída na rexión do Cáucaso era duns 20 000 animais e a densidade era de catro animais por quilómetro cadrado.