Based on studies of 16S-rRNA, they have been grouped into the subdivision of gram-positive eubacteria with high G:C content, with close phylogenetic relationship to Arthrobacter, Mycobacterium, Nocardia, and Streptomyces.
Os estudos do ARNr de 16S permitiron despois agrupalos na subdivisión de eubacterias grampositivas con alto contido G:C, cunha relación filoxenética moi próxima con Arthrobacter, Mycobacterium, Nocardia, e Streptomyces.