His 1937 work Genetics and the Origin of Species became a major influence on the synthesis and was awarded the US National Medal of Science in 1964, and the Franklin Medal in 1973.
O seu traballo de 1937 Genetics and the Origin of Species tivo unh grande influencia na síntese evolutiva e foi premiado coa Medalla Nacional da Ciencia de Estados Unidos de 1964, e coa Medalla Franklin en 1973.