Subgroups on the periphery of the central group are run by one dominant male, of a rank lower than the central males, and he maintains order in the group, and communicates messages between the central and peripheral males.
Os subgrupos na periferia do grupo central son dirixidos por un macho dominante, de rango inferior ao dos machos centrais, e el mantén a orde no grupo e comunica mensaxes entre os machos centrais e periféricos.