Intrinsic muscles of the foot originate and insert in the foot: the extensor digitorum brevis and extensor hallucis brevis on the dorsal aspect of the foot; all other intrinsic muscles—the dorsal and plantar interossei, flexor digiti minimi brevis, flexor hallucis brevis, flexor digitorum brevis, quadratus plantae (flexor accessorius), abductor digiti minimi, abductor hallucis, and lumbricals—are on the plantar side of the foot in the sole where they are organized into four layers.