The joint capsules are reinforced by medial and lateral collateral ligaments, and by plantar ligaments, which have grooves on their plantar surfaces for the long tendons of the digits (Fig. 6.109).
Four deep transverse metatarsal ligaments link the heads of the metatarsals together and enable the metatarsals to act as a single unified structure (Fig. 6.109). The ligaments blend with the plantar ligaments of the adjacent metatarsophalangeal joints.