France's David Casteu, riding a Yamaha, has won the fifth motorcycling stage of the Dakar Rally, a 136km timed run between Arequipa in Peru and Arica in Chile.

Compatriot Olivier Pain, was second on Wednesday's stage, 1 minute 09 seconds behind on his Yamaha and retained the overall lead.

Italy's Alessandro Botturi, on a Husqvarna, was third, 3:25 off the pace.

Pain has an overall lead of 1:15 on Casteu while defending champion Cyril Despres on a KTM, and who was fourth on Wednesday, is third overall, 6:07 behind.